Cloning and function analysis of Zea mays adverse stress inducible promoter
A promoter sequence, corn technology, applied in the field of bioengineering, can solve the problem of few inducible promoters, and achieve the effect of strong comprehensive resistance
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Embodiment 1
[0035]Embodiment 1: Cloning of the adversity stress inducible promoter of maize anion peroxidase gene (ZmapH)
[0036] The promoter of maize anion peroxidase gene (ZmapH) (promoter sequence comprises the DNA nucleotide sequence of -1bp to-1150bp region relative to the transcription initiation site of SEQ ID NO:1), in maize anion peroxidase The oxidase gene (ZmapH) was identified in the sequence of the 5' untranslated region.
[0037] The maize anion peroxidase gene (ZmapH) is registered in NCBI GenBank (accession number: AF037033.1), and the sequence listing shows the DNA sequence of the plant adversity stress-inducible promoter and 5' untranslated region of the above-mentioned gene of the present invention. exist figure 1 In , the start codon ATG for protein synthesis is underlined, and the base A of the transcription start site is shown with +1. And use the promoter analysis website to analyze the core elements of the promoter. Promoter analysis website http: / / www.dna.aff...
Embodiment 2
[0042] Embodiment 2: Construction of plant adversity stress inducible vector
[0043] The promoter of the maize anion peroxidase gene (ZmapH) cloned in Example 1 and the 5' untranslated region ZmapH Pro (see the sequence listing) of 543 bp were inserted into the vector to construct a plant stress-inducible vector.
[0044] Specifically, the plant expression vector pCAMBIA1301 and the recombinant plasmid pMD 18-T::ZmapH Pro were respectively digested with BamHI and NcoI, and then inserted into the BamHI and NcoI restriction sites of the vector pCAMBIA1301. The vector is called pCAMBIA 1301::ZmapH Pro, and it is used to drive the expression of GUS gene, which was identified by enzyme digestion ( Figure 5 ) obtained the promoter fragment, which was the same as the expected result.
[0045] exist Image 6 Among them, the gene GUS encoding β-glucuronidase was used as the reporter gene, and the selection marker was hygromycin resistance gene. In addition, 35s-pro represents the ...
Embodiment 3
[0046] Example 3: Identification of the activity of maize adversity stress-inducible promoters
[0047] By the heat shock transformation method, the vector pCAMBIA 1301::ZmapH Pro constructed in Example 2 is transferred to Agrobacterium tumefaciens EHA105, and the plasmid is extracted ( Figure 7 ) and PCR identification ( Figure 8 ).
[0048] In order to identify the stress-induced activity of the promoter, the method of Jefferson et al. (EMBO J, 1987) was used to test the activity of GUS after drought treatment, one of the stress treatments of mature embryos of maize.
[0049] More specifically, soak the corn seeds to accelerate germination, then cut the seeds in half longitudinally, induce culture with 20% PEG for 24 hours, put the corn seeds in the GUS detection solution at 37°C overnight, GUS detection solution: 1mg / ml X- gluc (5-bromo-4-chloro-3-indole-β-D-glucuronide), 50mM sodium phosphate buffer solution (PH=7.0), 10mM EDTA, 0.5mM potassium ferricyanide, 0.5mM ferr...
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