System and method for reducing intumescence injury of solanaceous plants
Supplemental UV radiation from an UV-LED light source with a peak wavelength of 300 to 322 nm addresses intumescence injury in solanaceous plants, improving growth and yield by mimicking optimal UV conditions.
Patent Information
- Application Number
- PCT/EP2024/083943
- Authority / Receiving Office
- WO · WO
- Patent Type
- Applications
- Current Assignee / Owner
- Priority Date
- 2023-12-04
- Filing Date
- 2024-11-28
- Publication Date
- 2025-06-12
AI Technical Summary
Solanaceous plants, such as tomato varieties, suffer from intumescence injury in environments deficient in solar UV radiation, leading to damage on leaves and stems, reduced plant growth, and decreased fruit yield.
The use of an UV-LED light source with a peak wavelength in the range of 300 to 322 nm provides supplemental UV radiation to solanaceous plants, effectively reducing intumescence injury by mimicking optimal UV radiation conditions.
The implementation of the UV-LED light source significantly reduces intumescence injury in solanaceous plants, enhancing plant growth, fruit yield, and overall health by providing the necessary UV radiation.
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Abstract
Description
SYSTEM AND METHOD FOR REDUCING INTUMESCENCE INJURY OF SOLANACEOUS PLANTSFIELD OF THE INVENTION[1] This invention relates to the field of plant cultivation. More specifically, the present invention provides an agricultural system for cultivating solanaceous plants comprising a plant cultivation area which is deficient in solar UV radiation, wherein said system comprises an UV-LED light source. The present invention further provides method for cultivating solanaceous plants in a controlled plant cultivation environment deficient in solar UV radiation, said method comprises subjecting the plant to supplemental UV radiation emitted by an UV-LED light source. The present invention further provides the use of supplemental UV radiation emitted by an UV-LED light source for reducing intumescence injury of solanaceous plants cultivated in a plant cultivation area which is deficient in solar UV radiation.BACKGROUND[2] Solanaceous plants can develop a type of damage on the leaves and stems called intumescence. Intumescence injury is characterized by the formation of galls and callus, and in extreme cases results in leaf death. Microscopic analysis of leaves of injured plants showed that spots with intumescence had defects in the outer waxy layer called the cuticle, and enlarged individual cells in the epidermis and other tissues that can ultimately rupture or collapse (Suzuki et al. (2020) Hort J 89: 597-574). The cuticle is a water-repellent layer that prevents water loss from the epidermis and protects against UV radiation and pathogen infection. In spots with intumescence this water repellence was lost and leaves damaged by intumescence had lower water content than normal leaves (Suzuki et al. (2020) Loc. cit.).[3] Conditions with high UV radiation and low relative humidity stimulate the development of the cuticle. By contrast, the cuticle does not develop well at high humidity and in the absence of UV, conditions that also lead to intumescence (Kubota et al. (2017) Sci Hortic 226: 366-371 ; Suzuki et al. (2020) Loc. cit.). In sensitive tomato varieties like Competition, intumescence severely injures plants in the absence of UV light. Indoor cultivation using LED lighting is particularly challenging because there is no UV radiation at all unless it is supplied by additional UV lamps. Also in glass greenhouses intumescence can occur because glass mostly blocks UV light and in winter UV radiation from the sun will be limited.[4] In previous studies, a broad range of UV wavelengths was applied, particularly by using gas discharge lights. Based on the use of such gas discharge lights emitting UV light, Kubota et al. (2017) Loc. cit. suggest that light in the UV-B spectrum is required to suppress intumescence. However, a more precise definition of the range of wavelengths is not provided, rendering the reported results not useful when the objective is to replace the gas discharge lights emitting UV light with LED light sources, which emit UV light in a much narrower wavelength band when compared to a gas discharge light source. Particularly, LED plant lighting systems conventionally comprise monochromatic LEDs that are characterized in that they emit light in very narrow bandwidth. Moreover, Kubota et al. (2017) Loc cit. fail to provide whether irradiation with light in the UV-A spectrum and / or the UV-C spectrum is required to effectively reduce intumescence injury in tomato plants.[5] It is an objective of the present invention to provide improved means and methods to reduce intumescence injury in solanaceous plants, which prevents subjecting said plants to UV light having a wavelength that is either ineffective or even is associated with negative effects on the cultivated solanaceous plants.SUMMARY OF INVENTION[6] The present invention relates to an agricultural system for cultivating solanaceous plants comprising a plant cultivation area which is deficient in solar UV radiation, wherein said system comprises an UV-LED light source having a peak wavelength in the range of 300 to 322 nm.[7] In addition, the present invention relates to a method for cultivating solanaceous plants in a controlled plant cultivation environment deficient in solar UV radiation, said method comprises subjecting the plant to supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of 300 to 322 nm.[8] In addition, the present invention relates to the use of supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of 300 to 322 nm for reducing intumescence injury of solanaceous plants cultivated in a plant cultivation area which is deficient in solar UV radiation.BRIEF DESCRIPTION OF THE FIGURES[9] Figure 1 : normalized non-UV-LED light spectrum
[0010] Figure 2: Normalized light spectrum of UV LEDs of different wavelengths used in the Examples. The region < 283 nm of the spectrum of 285 nm LEDs was mirrored from the region > 283 nm because it was out of range of the spectrometer. The measured peak wavelength was 283 nm and it is normal that there is some deviation from factory specified peak wavelength (285 nm in this case). It was assumed that the spectrum was symmetrical around the peak wavelength.
[0011] Figure 3. Seedlings 17 days after sowing (das) and 12 days after start of UV treatments. Treatment codes: UV intensity in pmol / m2 / s is given, followed by the wavelength.
[0012] Figure 4: Lower (abaxial) side of a leaf 24 days after sowing (das). Treatment codes: UV intensity in pmol / m2 / s is given, followed by the wavelength. The 285-nm UV treatment is missing because it was stopped after the seedling stage.
[0013] Figure 5: Effect of UV wavelength on tomato variety Competition at final harvest 34 das. The 285-nm UV treatment is missing because it was stopped after the seedling stage.
[0014] Figure 6: Effect of UV wavelength on tomato variety Competition at final harvest 34 days after sowing. The 285-nm UV treatment is missing because it was stopped after the seedling stage.
[0015] Figure 7: Effect of UV wavelength on a fully elongated leaf at the top of the plants tomato variety Competition at final harvest 34 days after sowing.
[0016] Figure 8: Effect of UV wavelength on the base of the stem of tomato variety Competition at final harvest 34 days after sowing.
[0017] Figure 9: Effect of UV wavelength on plant height of tomato variety Competition at final harvest 34 days after sowing. Data are means of 18 plants (0.12 308nm treatment) or 24 plants (Ctrl and 0.12 325nm treatment), error bars are standard deviation. Means with different letters are significantly different (Tukey test, P < 0.05).
[0018] Figure 10: Effect of UV wavelength on fresh weight of leaves and stem of tomato variety Competition at final harvest 34 days after sowing. Data are means of 18 plants (0.12 308nm treatment) or 24 plants (Ctrl and 0.12 325nm treatment), error bars are standard deviation. Means with different letters are significantly different (Tukey test, P < 0.05), lowercase: FW leaves, uppercase FW stem.
[0019] Figure 11 : Effect of intensity of 308-nm UV light on tomato variety Competition at final harvest 34 days after sowing.
[0020] Figure 12: Effect of intensity of 308-nm UV light on tomato variety Competition at final harvest 34 days after sowing.
[0021] Figure 13: Effect of intensity of 308-nm UV light on tomato variety Competition at final harvest 34 days after sowing.
[0022] Figure 14: Effect of intensity of 308-nm UV light on tomato variety Competition at final harvest 34 days after sowing.
[0023] Figure 15: Effect of intensity of 308-nm UV light on plant height of tomato variety Competition at final harvest 34 days after sowing. Data are means of 16 plants, except for the control (22 plants), error bars are standard deviation. Means with different letters are significantly different (Tukey test, P < 0.05).
[0024] Figure 16: Effect of intensity of 308-nm UV light on fresh weight (FW) of leaves and stem of tomato variety Competition at final harvest 34 days after sowing. Data are means of 16 plants, except for the control (22 plants), error bars are standard deviation. Means with different letters are significantly different (Tukey test, P < 0.05), lowercase: FW leaves, uppercase FW stem.
[0025] Figure 17: Effect of intensity of 308-nm UV light on tomato variety Competition at final harvest 31 days after sowing.
[0026] Figure 18: Effect of intensity of 308-nm UV light on tomato variety Competition at final harvest 31 days after sowing.
[0027] Figure 19: Effect of intensity of 308-nm UV light on a fully elongated leaf at the top of the plants of tomato variety Competition at final harvest 31 days after sowing.
[0028] Figure 20: Effect of intensity of 308-nm UV light on the base of the stem of tomato variety Competition at final harvest 31 days after sowing.
[0029] Figure 21 : Effect of intensity of 308-nm UV light on plant height of tomato variety Competition at final harvest 31 days after sowing. Data are means of 15 plants; error bars are standard deviation. Means with different letters are significantly different (Tukey test, P < 0.05).
[0030] Figure 22: Effect of intensity of 308-nm UV light on fresh weight (FW) of leaves and stem of tomato variety Competition at final harvest 31 days after sowing. Data are means of 15 plants; error bars are standard deviation. Means with different letters are significantly different (Tukey test, P < 0.05), lowercase: FW leaves, uppercase FW stem.DETAILED DESCRIPTION OF THE INVENTIONGeneral definitions
[0031] It is to be understood that this invention is not limited to the particular methodology or protocols. It is also to be understood that the terminology used herein is for the purpose of describing particular embodiments only, and is not intended to limit the scope of the present invention which will be limited only by the appended claims. It must be noted that as used herein and in the appended claims, the singular forms "a," "and," and "the" include plural reference unless the context clearly dictates otherwise. Thus, for example, reference to "a light source" is a reference to one or more light sources and includes equivalents thereof known to those skilled in the art, and so forth. The term "about" is used herein to mean approximately, roughly, around, or in the region of. When the term "about" is used in conjunction with a numerical range, it modifies that range by extending the boundaries above and below the numerical values set forth. In general, the term "about" is used herein to modify a numerical value above and below the stated value by a variance of 20 percent, preferably 10 percent up or down (higher or lower). As used herein, the word "or" means any one member of a particular list and also includes any combination of members of that list. The words "comprise," "comprising," "include," "including," and "includes" when used in this specification and in the following claims are intended to specify the presence of one or more stated features, integers, components, or steps, but they do not preclude the presence or addition of one or more other features, integers, components, steps, or groups thereof. For clarity, certain terms used in the specification are defined and used as follows:
[0032] As used herein, the term "agricultural system for cultivating solanaceous plants" refers to any arrangement of equipment that is suitable for cultivating solanaceous plants. Said agricultural system for cultivating solanaceous plants accordingly preferably comprises a plurality of solanaceous plants, more preferably solanaceous plants susceptible to intumescence injury, in any stage of plant development. The agricultural system according to the present invention comprises a plant cultivation area, which refers to the part of the agricultural system which provides all means required for cultivating solanaceous plants, particularly a plant cultivation substrate. Accordingly, said plant cultivation substrate provides a suitable support for the plant root system. The term “agricultural system for cultivating solanaceous plants” according to the present invention accordingly comprises, but is not limited to a tunnel, a greenhouse, such as a glass greenhouse, a climate chamber and an indoor farming system.
[0033] The term “deficient in solar UV radiation” in the context of the present invention means that the solar UV radiation in the plant cultivation area is such that it causes adverse effects on the cultivated plants, such as intumescence injury, when compared to plants that are cultivated underoptimal UV radiation conditions. Such deficiency in solar UV radiation preferably comprises an insufficient intensity of the UV radiation and / or an insufficient wavelength composition of the UV radiation. Adverse effects caused by a deficiency in solar UV radiation include, but are not limited to, damage on the leaves and / or stems, formation of galls and / or callus, reduced plant growth (expressed in fresh weight), reduced fruit yield and delayed fruit production.
[0034] The term “means suitable for controlling lighting” or “lighting system” in the context of the present invention refers to a subassembly of an agricultural system suitable for cultivating plants comprising one or more artificial light sources and which is capable of emitting light which spectral composition and intensity is suitable for cultivating plants, particularly solanaceous plants. The light emitted by the lighting system according to the present invention comprises a UV-LED light source and optionally an artificial non-UV light source.
[0035] The term “UV light” or “UV radiation” or “UV” refers to a range of light at the ultraviolet end of the visible spectrum. Preferably, “UV light” is light having a wavelength in a range of 100 to 400 nm. The term "peak wavelength in a range" accordingly may also include light emitters that emit light outside the indicated range in addition to the peak emission at the indicated wavelength range. Different UV bands are defined as UV-C, UV-B, and UV-A. Preferably, “UV-C” is light having a wavelength in a range of 100 to 280 nm. Preferably, “UV-B” is light having a wavelength in a range of more than 280 to 315 nm. Preferably, “UV-A” is light having a wavelength in a range of more than 315 to 400 nm.
[0036] The term “supplemental UV light" or “supplemental UV radiation" as used herein refers any to UV light emitted by an artificial UV light source, both in addition to the solar UV radiation that may already be present in the plant cultivation areas (like in a glass green house) as well as in a plant cultivation area in which there is essentially no solar UV radiation (like in a climate chamber or an indoor farming system).
[0037] The term “far-red light” refers to a range of light at the extreme red end of the visible spectrum. Preferably, “far-red light” is light having a wavelength in a range of 700 to 800 nm. The term "supplemental far-red light" refers to far-red light that is applied, e.g. to a plant, in addition to the already present ambient light such as the available sunlight and / or artificial light.
[0038] The term "artificial light source" refers to any man-made light source that is emitting light. Examples of such artificial light source include incandescent light sources, gas discharge light sources and LED light sources. An "incandescent light source" is an electric light source comprising a wire filament to which an electric current is applied causing the filament to glow, thereby emitting light over a broad wavelength spectrum. Incandescent light sources have several disadvantages and accordingly are not commonly used in horticulture. The term "gas discharge light source" refers to an artificial light source that generate light over a relatively broad wavelength spectrum by sending an electric discharge through an ionized gas. Gas discharge light sources useful for irradiation of plants in horticulture are well-known in the art. The term "LED light source" relates to a light source that wherein Light Emitting Diodes are used for illumination. LED light sources, particularly monochromatic LED light sources, are characterized in that they emit light in a specific band of wavelengths which is much narrower when compared to light emitted by other artificial light sources. LED light sources useful for irradiation of plants in horticulture are well-known in the art. Such LED light sources are typically capable of emitting a Blue lightcomponent (B), a Green light component (G), a Red light component (R) and a Far-red light component (Fr), wherein the different light components are emitted by a plurality of narrow bandwidth LED light sources. The term “narrow bandwidth” in relation to a LED light source has a well- recognized meaning and accordingly refers to an LED light source characterized by emitting light with in a specific and limited range of wavelengths. In the context of the present invention, the term “narrow bandwidth LED light source” preferably refers to a LED light source having a full width at half maximum (FWHM) emission spectrum of at most 30 nm, more preferably of at most 29 nm, 28 nm, 27 nm, 26 nm, 25 nm, 24 nm, 23 nm, 22 nm, 21 nm, 20 nm, 18 nm, 16 nm, 15 nm, 14 nm, 13 nm, 12 nm, or 11 nm, most preferably of at most 10 nm. Determining the FWHM value of a LED light source is very well-known in the art and represent the range of wavelengths where the intensity of the light emitted by the LED light source is at least 50% of the intensity at the peak wavelength of said LED light source.
[0039] As used herein the term "artificial UV light source" refers to an artificial light source as defined herein that is capable of emitting UV light as defined herein. As used herein the term "UV- LED light source" refers to a LED light source capable of emitting UV light as defined herein. Accordingly, a “narrow bandwidth UV-LED light source" refers to a LED light source capable of emitting UV light in a specific and limited range of wavelengths. In the context of the present invention, the term “narrow bandwidth UV-LED light source” preferably refers to a UV-LED light source having a full width at half maximum (FWHM) emission spectrum of at most 25 nm, more preferably of at most 20 nm, 18 nm, 16 nm, 15 nm, 14 nm, 13 nm, 12 nm, or 11 nm, most preferably of at most 10 nm.
[0040] The term "non-UV light source" refers to an artificial light source that emits “non-UV light”, which in the context of the present invention refers to light in the visible spectrum having a wavelength of more than 400 nm. Preferably, the non-UV light source emits light having a wavelength of more than 400 nm up to 800nm. The term “supplemental non-UV light" or “supplemental non- UV radiation" as used herein refers any to non-UV light emitted by an artificial non-UV light source, both in addition to the solar non-UV radiation that may already be present in the plant cultivation areas (like in a glass green house) as well as in a plant cultivation area in which there is essentially no solar non-UV radiation (like in a climate chamber or an indoor farming system).
[0041] The term "phytochrome stationary state (PSS) value" refers to the ratio of the concentration of the Prisoform of phytochrome to the total concentration of both isoforms under constant irradiation by a light source (Sager et al. (1988) Trans. ASAE31 , 1882-1889. doi: 10.13031 / 2013.30952). Phytochrome exists in two states, or isoforms. In its ground state (identified as Pr), phytochrome strongly absorbs red light, and so appears turquoise-blue in concentrated solution in vitro. When it absorbs a red photon, however, it changes its physical shape to form its physiologically active state Pfr. In doing so, its peak spectral absorptance shifts towards the far-red, with a concentrated solution of phytochrome appearing more greenish in colour. When phytochrome is in its Pfr state, it may absorb a far-red photon and change once again into its Prstate. This bi-stable behaviour makes phytochrome an ideal biochemical switch, with the Prisoform serving as the signalling state to the plant.
[0042] The term "intumescence injury" or "intumescence" as used herein refers to a physiological disorder characterized by hypertrophy and possibly hyperplasia of plant cells. In susceptible sol- anaceous plants is typically observed in the leaves and stems which may lead to the formation ofgalls and callus, and in extreme cases results in leaf death. Microscopic analysis of plant leaves showed that spots with intumescence injury have defects in the outer waxy layer called the cuticle, and enlarged individual cells in the epidermis and other tissues that can ultimately rupture or collapse; see e.g. Suzuki et al. (2020) Loc. cit. The cuticle is a water-repellent layer that prevents water loss from the epidermis and protects against UV radiation and pathogen infection. In spots with intumescence this water repellence was lost and leaves damaged by intumescence had lower water content than normal leaves. The level of sensitivity to intumescence injury of a given solanaceous variety can be easily determined by comparing plants of said variety that are cultivated under conditions which are deficient in UV radiation with plants that are cultivated under conditions comprising sufficient UV radiation.
[0043] The term “means suitable for controlling temperature” or “temperature control system” in the context of the present invention refers to a subassembly of an agricultural system suitable for cultivating plants comprising one or more means which are able and adapted to control the ambient temperature of the cultivation space. The term “ambient temperature of the cultivation space” refers to the temperature of the atmosphere of the plant cultivation space. Optionally, temperature control system further comprises one or more means which are able and adapted to control the root temperature of the plant root system, and / or one or more means which are able and adapted to control the leaf temperature of the leaf system, wherein said root temperature and / or leaf temper able and adapted to root temperature and / or leaf temperature that differs from an ambient temperature. Particularly, the temperature control system may comprise heating means and / or cooling means.
[0044] The term “means suitable for controlling humidity” or “humidity control system” in the context of the present invention refers to a subassembly of an agricultural system suitable for cultivating plants comprising one or more means which are able and adapted to control the humidity of the atmosphere of the plant cultivation space.
[0045] The term “means suitable for controlling carbon dioxide (CO2) content” or “carbon dioxide control system” in the context of the present invention refers to a subassembly of an agricultural system suitable for cultivating plants comprising one or more means which are able and adapted to control the controlling carbon dioxide content of the atmosphere of the plant cultivation space.
[0046] The term “means suitable for controlling shading” or “shading system” in the context of the present invention refers to a subassembly of an agricultural system suitable for cultivating plants comprising one or more means which are able and adapted to control the intensity of solar radiation in the plant cultivation space.
[0047] The term “means suitable for controlling ventilation” or “ventilation system” in the context of the present invention refers to a subassembly of an agricultural system suitable for cultivating plants comprising one or more means which are able and adapted to control the air movement in the plant cultivation space.
[0048] The term “means suitable for controlling irrigation” or “irrigation system” in the context of the present invention refers to a subassembly of an agricultural system suitable for cultivating plants comprising one or more means which are able and adapted to control the distribution of water provided to the cultivated plants. The irrigation system preferably comprises a drip irrigationsystem. The term “means suitable for controlling nutrient distribution” or “nutrient distribution system” in the context of the present invention refers to a subassembly of an agricultural system suitable for cultivating plants comprising one or more means which are able and adapted to control the distribution of plant nutrients provided to the cultivated plants.
[0049] "Solanaceous plants" or "plants of the family Solanaceae" are plants of the botanical family Solanaceae, i.e. any plant of the family Solanaceae, including wild solanaceous plants and cultivated solanaceous plants. The botanical family Solanaceae consists about 98 genera of which the genera Solanum and Capsicum are the commercially most relevant as they comprise many domesticated species that are widely cultivated and used as food crops with high economic importance. In the context of the present invention, particularly relevant solanaceous plants are tomato plants, eggplants, potato plants, pepper plants, or tobacco plants.
[0050] The genus Solanum consists of about 1330 species, including the highly important food crops tomato plants, eggplant or aubergine plants and potato plants.
[0051] Tomato plants are herbaceous plants of the family Solanaceae that are of particular relevance in the context of the present invention. In the context of the present invention, the term “tomato plants” refers to Solanum lycopersicum plants and wild relatives of Solanum lycopersi- cum. Tomato plants are perennial in their native habitat but cultivated as an annual. Cultivated tomato plants typically grow to 1-3 meters (3-10 ft) in height. Tomato fruits are botanically berrytype fruits, they are considered culinary vegetables. Fruit size varies according to cultivar, with a width range of about 1-10 cm (about 0.5-4 inches). Solanum lycopersicum is also known as Ly- copersicon lycopersicum (L.) H. Karst, or Lycopersicon esculentum Mill. The term "cultivated tomato plant" or "cultivated tomato" refers to plants of Solanum lycopersicum, e.g. varieties, breeding lines or cultivars of the species S. lycopersicum, cultivated by humans and having good agronomic characteristics. "Wild relatives of Solanum lycopersicum " include S. arcanum, S. chmielewskii, S. neorickii (= L. parviflorum), S. cheesmaniae, S. galapagense, S. pimpinellifolium, S. chilense, S. corneliomulleri, S. habrochaites (= L. hirsutum), S. huaylasense, S. sisymbriifo- lium, S. peruvianum, S. hirsutum or S. pennellii. Tomato plants are diploid and have 12 pairs of homologous chromosomes, numbered 1 to 12. Tomato plants may have a varying sensitivity to intumescence injury. Particularly Solanum habrochaites, Solanum pimpinelifolium and tomato plants used as a rootstock are sensitivity to intumescence injury.
[0052] "Eggplant" or "aubergine plants" or “Solanum melongena plants” are further herbaceous plants of the family Solanaceae that are of particular relevance in the context of the present invention. The egg-shaped, glossy, dark purple to white fruit has white flesh with a meaty texture. S. melongena plants grow about 40-150 cm (about 1.3-5 ft) tall, with large, coarsely lobed leaves that are about 10-20 cm (about 3-8 in) long and about 5-10 cm (about 2-4 in) broad.
[0053] The term "potato" or "potato plants" refers to any Solanum lycopersicum plants or wild relatives thereof, which are characterized by its starchy tubers growing underground.
[0054] The term “pepper plants” as used herein refers to plants of domesticated Capsicum plants, namely C. annuum, C. baccatum, C. chinense, C. frutescens, and C. pubescens. Fruits of Capsicum plants, often named as "peppers" or "pepper fruits", can vary tremendously in colour, shape, and size both between and within species.
[0055] Capsicum annuum L. plants are herbaceous plants of the family Solanaceae that are of particular relevance in the context of the present invention. Capsicum annuum plants reach about 0.5-1.5 meters (about 20-60 inches). Single white flowers bear the pepper fruit which is green when unripe, changing principally to red, although some varieties may ripen to brown or purple. While the species can tolerate most climates, they are especially productive in warm and dry climates. Cultivated plants of the species Capsicum annuum include different types of peppers, such as bell peppers, cayenne peppers, paprika, and jalapenos. "Capsicum annuum chromosome 3" refer to the Capsicum annuum chromosome 3, as known in the art (see Capsicum annuum cv CM334 genome chromosomes (release 1.55) and Capsicum annuum LICD10X genome chromosomes (v1.0) and Capsicum annuum zunla genome chromosomes (v2.0).
[0056] The term “Tobacco plants” refers to plants in the genus Nicotiana of the family Solanaceae, of which the cured leaves are used as a stimulant. More than 70 species of tobacco are known, of which the main commercially cultivated species are N. tabacum and N. rustica.
[0057] As used herein, the term "plant" preferably includes the whole plant, including different developmental stages, such as seedlings, immature and mature, etc. When “seeds of a plant” are referred to, these either refer to seeds from which the plant can be grown or to seeds produced on the plant, after fertilization. Which meaning is used is evident from the context.
[0058] "Plant variety" is a group of plants within the same botanical taxon of the lowest grade known, which (irrespective of whether the conditions for the recognition of plant breeder’s rights are fulfilled or not) can be defined on the basis of the expression of characteristics that result from a certain genotype or a combination of genotypes, can be distinguished from any other group of plants by the expression of at least one of those characteristics, and can be regarded as an entity, because it can be multiplied without any change. Therefore, the term “plant variety” cannot be used to denote a group of plants, even if they are of the same kind, if they are all characterized by the presence of one or two loci or genes (or phenotypic characteristics due to these specific loci or genes), but which can otherwise differ from one another enormously as regards the other loci or genes.
[0059] The term "cultivar" (or “cultivated” plant) is used herein to denote a plant having a biological status other than a "wild" status, which "wild" status indicates the original non-cultivated, nondomesticated, or natural state of a plant or accession, and the term cultivated does not include such wild, or weedy plants. The term cultivar does include material with good agronomic characteristics, such as breeding material, research material, breeding lines, elite breeding lines, synthetic population, hybrid, founder stock / base population, inbred lines, cultivars (open pollinated of hybrid cultivar), segregating population, mutant / genetic stock, and advanced / improved cultivar. The so-called heirloom varieties or cultivars, i.e. open pollinated varieties or cultivars commonly grown during earlier periods in human history and often adapted to specific geographic regions, are in one aspect of the invention encompassed herein as cultivated plants. In one embodiment the term cultivar also includes landraces, i.e. plants (or populations) selected and cultivated locally by humans over many years and adapted to a specific geographic environment and sharing a common gene pool.
[0060] The term "food" is any substance consumed to provide nutritional support for the body. It is usually of plant or animal origin, and contains essential nutrients, such as carbohydrates, fats,proteins, vitamins, or minerals. The substance is ingested by an organism and assimilated by the organism's cells in an effort to produce energy, maintain life, or stimulate growth. The term food includes both substance consumed to provide nutritional support for the human and animal body.
[0061] The term "seed production plant" refers to a plant that is specifically cultivated with the purpose of producing plant seeds for subsequent sowing. Accordingly, the produce of a seed production plant is preferably not used as food.
[0062] A "plant line" or "breeding line" refers to a plant and its progeny. As used herein, the term "inbred line" refers to a plant line which has been repeatedly selfed and is nearly homozygous. Thus, an “inbred line” or “parent line” refers to a plant which has undergone several generations (e.g. at least 4, 5, 6, 7 or more) of inbreeding, resulting in a plant line with a high uniformity.
[0063] "Selfing" refers to self-pollination of a plant, i.e., the transfer of pollen from the anther to the stigma of the same plant.
[0064] "Crossing" or cross-pollination refers to the mating of two parent plants, e.g. pollinating a female parent line with pollen of a male parent line.
[0065] "F1 , F2, F3, etc. " refers to the consecutive related generations following a cross between two parent plants or parent lines. The plants grown from the seeds produced by crossing two plants or lines is called the F1 generation. Selfing the F1 plants results in the F2 generation, etc.
[0066] A "female plant" is to be understood herein as a plant that is pollinated by a male parent.
[0067] A "male plant" is to be understood herein as a plant of which the pollen is used to pollinate a female plant.
[0068] "F1 hybrid" plant (or "F1 hybrid seed") is the generation obtained from crossing two inbred parent lines. Thus, F1 hybrid seeds are seeds from which F1 hybrid plants grow. F1 hybrids are more vigorous and higher yielding, due to heterosis. Inbred lines are essentially homozygous at most loci in the genome.
[0069] "Pollination" is the process by which pollen is transferred to the stigma of the flower, thereby enabling fertilization and reproduction.
[0070] "Emasculation" is understood herein as the removal of the anthers to prevent self-pollination.
[0071] "Sowing" refers to the process of placing the seed in an appropriate medium for germination, so that it is able to grow into a plant. The medium can e.g. encompass soil, various types of rockwool, (liquid) culture media, etc.
[0072] The term "fruit" refers to the seed-bearing structure in flowering plants.
[0073] "Harvest" is understood herein to be the collection of fruits and / or seeds developed in the fruits.
[0074] "Average" refers herein to the arithmetic mean.
[0075] It is understood that comparisons between different cultivation conditions involves cultivating a number of plants of a line (or variety) (e.g. at least 5 plants, preferably at least 10 plants per line) under the
[0076] relevant test conditions and control conditions and the determination of differences, preferably statistically significant differences, between the plants when grown under the relevant cultivation conditions. Preferably the plants are of the same line or variety.Agricultural system for cultivating solanaceous plants
[0077] The present invention provides an agricultural system for cultivating solanaceous plants comprising a plant cultivation area which is deficient in solar UV radiation, wherein said system comprises an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm.
[0078] The inventors found that the use of supplemental UV irradiation provided by an UV-LED light source having a peak wavelength in the range of 300 to 322 nm is sufficient to reduce or even prevent intumescence injury when cultivating solanaceous plants, including tomato plant varieties which are particularly sensitive to intumescence injury. It was surprisingly found that supplemental UV light having a wavelength other than the range of 300 to 322 nm not only is ineffective to prevent intumescence injury, but even may have a negative effect on plant health. This is surprising because it was conventionally believed that also UV-A and UV-C is required to prevent intumescence injury in tomato plants grown under UV deficient cultivation conditions. Currently UV-LEDs are not available for commercial glass greenhouse application.
[0079] The present invention defines the UV wavelength significantly more precisely than earlier. An earlier study concluded that UV-B (280-315 nm) is required to suppress intumescence (Kubota et al. (2017) Loc. cit.), but the UV light source as used in this study had a broad range of roughly 300-400 nm and a peak around 345 nm. Within this range, we have shown that a peak wavelength of 285 nm, still in the UV-B range, is highly damaging to tomato plants, while a peak wavelength of 322 or more is not sufficiently effective in suppressing intumescence. These findings also put into question the previously reported effective intensity or daily dose, because only a fraction of the wavelengths contribute effectively to suppressing intumescence.
[0080] The agricultural system according to the present invention accordingly is suitable for the cultivation of solanaceous plants. Preferably, the solanaceous plants cultivated in the agricultural system according to the present invention are potato plants, tobacco plants, pepper plants, eggplant or tomato plants. Most preferably, the solanaceous plants cultivated in the agricultural system according to the present invention are tomato plants.
[0081] The agricultural system for cultivating solanaceous plants according to the present invention accordingly comprises a plant cultivation area which is deficient in solar UV radiation. Typical float glass used in glass greenhouses transmits very little light of wavelengths shorter than 400 nm, so in the UV range, and transmittance is near zero around 310 nm, almost exactly the wavelength most effective in suppressing intumescence. Hence for natural sunlight to suppress intumescence in glass greenhouses, intensity needs to be high to reach a sufficient UV-B intensity in the desired wavelength bandwidth. That means in glass greenhouses in winter under low light and in indoor systems, supplemental UV-B light is essential to cultivate sensitive varieties without severe intumescence injury.
[0082] The agricultural system for cultivating solanaceous plants according to the present invention accordingly comprises an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm. Preferably, the agricultural system for cultivating solanaceous plants according to the present invention comprises an LIV-LED light source having a peak wavelength in the range of 300 to 320 nm, 300 to 318 nm, 300 to 316 nm, 300 to 315 nm, 300 to 314 nm, 300 to 313 nm,300 to 312 nm, 300 to 311 nm, 300 to 310 nm, 301 to 322 nm, 301 to 320 nm, 301 to 318 nm,301 to 316 nm, 301 to 315 nm, 301 to 314 nm, 301 to 313 nm, 301 to 312 nm, 301 to 311 nm,301 to 310 nm, 302 to 322 nm, 302 to 320 nm, 302 to 318 nm, 302 to 316 nm, 302 to 315 nm,302 to 314 nm, 302 to 313 nm, 302 to 312 nm, 302 to 311 nm, 302 to 310 nm, 303 to 322 nm,303 to 320 nm, 303 to 318 nm, 303 to 316 nm, 303 to 315 nm, 303 to 314 nm, 303 to 313 nm,303 to 312 nm, 303 to 311 nm, 303 to 310 nm, 304 to 322 nm, 304 to 320 nm, 304 to 318 nm,304 to 316 nm, 304 to 315 nm, 304 to 314 nm, 304 to 313 nm, 304 to 312 nm, 304 to 311 nm,304 to 310 nm, 305 to 322 nm, 305 to 320 nm, 305 to 318 nm, 305 to 316 nm, 305 to 315 nm,305 to 314 nm, 305 to 313 nm, 305 to 312 nm, 305 to 311 nm, 305 to 310 nm, 306 to 322 nm,306 to 320 nm, 306 to 318 nm, 306 to 316 nm, 306 to 315 nm, 306 to 314 nm, 306 to 313 nm,306 to 312 nm, 306 to 311 nm or 306 to 310 nm. More preferably, the agricultural system for cultivating solanaceous plants according to the present invention accordingly comprises an UV- LED light source having a peak wavelength in the range of 305 to 315 nm.
[0083] Preferably, the agricultural system for cultivating solanaceous plants according to the present invention does not comprise a gas discharge UV light source. More preferably, the agricultural system for cultivating solanaceous plants according to the present invention does not comprise other supplemental UV light sources than the UV-LED light source according to the present invention.
[0084] The application of supplemental UV radiation in a work environment also carries a risk because the UV light is damaging to the eyes and skin and direct exposure should be avoided. This means that preferably no supplemental UV radiation is applied at moments when there are humans present in the plant growth space. Preferably, the the agricultural system for cultivating solanaceous plants according to the present invention comprises means to reduce or prevent that humans present in the plant growth space are irradiated with the supplemental UV light. Such means may include means that allow switching off the UV-LED light source when humans are present in the plant growth space, e.g. by using an automated system like a clock switch or a motion detector switch.
[0085] In one aspect, the UV-LED light source comprised in the system according to present invention is a narrow bandwidth UV-LED light source having a full width at half maximum (FWHM) emission spectrum of at most 30 nm. Preferably, the UV-LED light source comprised in the system according to present invention is a narrow bandwidth UV-LED light source having a full width at half maximum (FWHM) emission spectrum of at most 29 nm, 28 nm, 27 nm, 26 nm, 25 nm, 24 nm, 23 nm, 22 nm, 21 nm, 20 nm, 19 nm, 18 nm, 17 nm, 16 nm, 15 nm, 14 nm, 13 nm, 12 nm, 11 nm or 10 nm.
[0086] Preferably, the agricultural system for cultivating solanaceous plants according to the present invention comprises an UV-LED light source having a peak wavelength in the range of (i.e. a peak wavelength of) 300 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peakwavelength of 300 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 30nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 30 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 30 nm.
[0087] More preferably, the agricultural system for cultivating solanaceous plants according to the present invention comprises an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 311nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 27 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 27 nm.
[0088] Even more preferably, the agricultural system for cultivating solanaceous plants according to the present invention comprises an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 315 nm and a FWHMemission spectrum of at most 25 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 315nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 25 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 25 nm.
[0089] Particularly preferably, the agricultural system for cultivating solanaceous plants according to the present invention comprises an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 322 nm and a FWHMemission spectrum of at most 22 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 22 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 22 nm.
[0090] More particularly preferably, the agricultural system for cultivating solanaceous plants according to the present invention comprises an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 313 nm and a FWHM emissionspectrum of at most 21 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 313 nm anda FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 21 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 21 nm.
[0091] Even more particularly preferably, the agricultural system for cultivating solanaceous plants according to the present invention comprises an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM spectrum of at most 20 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 318 nm and a FWHM emissionspectrum of at most 20 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 20 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 20 nm.
[0092] Most preferably, the agricultural system for cultivating solanaceous plants according to the present invention comprises an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM spectrum of at most 18 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 18 nm, apeak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of atmost 18 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 18 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 18 nm.
[0093] The LIV-LED light source according to the present invention is capable of emitting an effective amount of UV radiation during the time period that the plants in the plant cultivation area are subjected to supplemental UV radiation (also indicated herein as “UV photoperiod”). An effective amount of UV radiation is characterized in that the UV-LED light source is capable of obtaining an intensity of UV light at plant height that is sufficient to compensate the deficiency in solar UV radiation, particularly that is sufficient to prevent that the adverse effects of the deficiency in solar UV radiations are reduced or even are fully absent. In one aspect, the UV-LED light source comprised in the system according to the present invention is capable of emitting UV radiation having an intensity of 0.1 to 2 pmol / m2 / s at plant height. In the context of the present invention, it was surprisingly found that intumescence injury in solanaceous plants could be further improved when 308-nm UV light intensity doubled from 0.12 pmol / m2 / s to 0.24 pmol / m2 / s or even to 0.36 pmol / m2 / s.
[0094] Preferably, the UV-LED light source comprised in the system according to the present invention is capable of emitting UV radiation having an intensity at plant height of 0.1 to 2 pmol / m2 / s, 0.2 to 2 pmol / m2 / s, 0.3 to 2 pmol / m2 / s, 0.1 to 1.5 pmol / m2 / s, 0.2 to 1.5 pmol / m2 / s, 0.3 to 1.5 pmol / m2 / s, 0.1 to 1 pmol / m2 / s, 0.2 to 1 pmol / m2 / s or 0.3 to 1 pmol / m2 / s.
[0095] Accordingly, the agricultural system for cultivating solanaceous plants according to the present invention comprises an UV-LED light source having a peak wavelength in the range of (i.e. a peak wavelength of) 300 to 322 nm, a FWHM emission spectrum of at most (i.e. a FWHM spectrum of at most) 30 nm and is capable of emitting UV radiation having an intensity at plant height of (i.e. intensity of) 0.1 to 2 pmol / m2 / s. Preferably, the agricultural system for cultivating solanaceous plants according to the present invention comprises an UV-LED light source having: a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to320 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.2 to 1 .5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.2 to 1 .5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most27 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.3 to 1 pmol / m2 / s; or a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.3 to 1 pmol / m2 / s.
[0096] The agricultural system for cultivating solanaceous plants according to the present invention comprises means allowing irradiation of the plants comprised in the plant cultivation area with an effective amount of non-UV light. Said non-UV light may originate from the sun (e.g. in a greenhouse) and / or from an artificial light source. An effective amount of non-UV light is characterized in that the intensity and spectral composition of the non-UV light in the plant cultivation area is suitable for solanaceous plant cultivation. The agricultural system for cultivating solanaceous plants according to the present invention accordingly comprises a subassembly which allows irradiation of the cultivated solanaceous plants with non-UV light which spectral composition and intensity is suitable for cultivating solanaceous plants. Such non-UV light having a spectral composition and intensity useful in solanaceous plant cultivation is known in the art.
[0097] In one aspect, the system according to the present invention further comprises an artificial non-UV light source. Preferably, the artificial non-UV light source is a "non-UV-LED light source". In horticultural lighting non-UV LED-based light sources have several advantages including high energy efficiency and relatively low costs.
[0098] In one aspect, the non-UV light source comprised in the system of the present invention is capable of emitting non-UV light with an intensity at plant height of 5-1000 pmol / m2 / s. Preferably, the non-UV light source comprised in the system of the present invention is capable of emitting non-UV light having an intensity at plant height of 5-1000 pmol / m2 / s, 10-1000 pmol / m2 / s, 15-1000 pmol / m2 / s, 20-1000 pmol / m2 / s, 25-1000 pmol / m2 / s, 30-1000 pmol / m2 / s, 35-1000 pmol / m2 / s, 40- 1000 pmol / m2 / s, 50-1000 pmol / m2 / s, 60-1000 pmol / m2 / s, 70-1000 pmol / m2 / s, 80-1000 pmol / m2 / s,90-1000 pmol / m2 / s, 100-1000 pmol / m2 / s, 5-500 pmol / m2 / s, 10-500 pmol / m2 / s, 15-500 pmol / m2 / s,20-500 pmol / m2 / s, 25-500 pmol / m2 / s, 30-500 pmol / m2 / s, 35-500 pmol / m2 / s, 40-500 pmol / m2 / s, 50-500 pmol / m2 / s, 60-500 pmol / m2 / s, 70-500 pmol / m2 / s, 80-500 pmol / m2 / s, 90-500 pmol / m2 / s, 100-500 pmol / m2 / s, 5-250 pmol / m2 / s, 10-250 pmol / m2 / s, 15-250 pmol / m2 / s, 20-250 pmol / m2 / s, 25-250 pmol / m2 / s, 30-250 pmol / m2 / s, 35-250 pmol / m2 / s, 40-250 pmol / m2 / s, 50-250 pmol / m2 / s,60-250 pmol / m2 / s, 70-250 pmol / m2 / s, 80-250 pmol / m2 / s, 90-250 pmol / m2 / s, 100-250 pmol / m2 / s,5-200 pmol / m2 / s, 10-200 pmol / m2 / s, 15-200 pmol / m2 / s, 20-200 pmol / m2 / s, 25-200 pmol / m2 / s, 30- 200 pmol / m2 / s, 35-200 pmol / m2 / s, 40-200 pmol / m2 / s, 50-200 pmol / m2 / s, 60-200 pmol / m2 / s, 70- 200 pmol / m2 / s, 80-200 pmol / m2 / s, 90-200 pmol / m2 / s, 100-200 pmol / m2 / s, 5-180 pmol / m2 / s, 10-180 pmol / m2 / s, 15-180 pmol / m2 / s, 20-180 pmol / m2 / s, 25-180 pmol / m2 / s, 30-180 pmol / m2 / s, 35-180 pmol / m2 / s, 40-180 pmol / m2 / s, 50-180 pmol / m2 / s, 60-180 pmol / m2 / s, 70-180 pmol / m2 / s, 80-180 pmol / m2 / s, 90-180 pmol / m2 / s, 100-180 pmol / m2 / s, 5-170 pmol / m2 / s, 10-170 pmol / m2 / s, 15- 170 pmol / m2 / s, 20-170 pmol / m2 / s, 25-170 pmol / m2 / s, 30-170 pmol / m2 / s, 35-170 pmol / m2 / s, 40-170 pmol / m2 / s, 50-170 pmol / m2 / s, 60-170 pmol / m2 / s, 70-170 pmol / m2 / s, 80-170 pmol / m2 / s, 90-170 pmol / m2 / s, 100-170 pmol / m2 / s, 5-160 pmol / m2 / s, 10-160 pmol / m2 / s, 15-160 pmol / m2 / s, 20- 160 pmol / m2 / s, 25-160 pmol / m2 / s, 30-160 pmol / m2 / s, 35-160 pmol / m2 / s, 40-160 pmol / m2 / s, 50- 160 pmol / m2 / s, 60-160 pmol / m2 / s, 70-160 pmol / m2 / s, 80-160 pmol / m2 / s, 90-160 pmol / m2 / s, 100- 160 pmol / m2 / s, 5-150 pmol / m2 / s, 10-150 pmol / m2 / s, 15-150 pmol / m2 / s, 20-150 pmol / m2 / s, 25-150 pmol / m2 / s, 30-150 pmol / m2 / s, 35-150 pmol / m2 / s, 40-150 pmol / m2 / s, 50-150 pmol / m2 / s, 60-150 pmol / m2 / s, 70-150 pmol / m2 / s, 80-150 pmol / m2 / s, 90-150 pmol / m2 / s, or 100-150 pmol / m2 / s.
[0099] In one aspect, the non-UV light source comprised in the system of the present invention is capable of emitting non-UV light comprising: a Blue light component (B) having a peak wavelength of more than 400 nm to at most 495 nm (preferably 435-485 nm), a Green light component (G) having a peak wavelength of more than 495 nm to at most 590 nm, (preferably 500-550 nm), a Red light component (R) having a peak wavelength of more than 590 nm to at most 700 nm (preferably 600-650 nm) and a Far-red light component (Fr) having a peak wavelength of more than 700 to at most 800 nm. The spectral composition of the non-UV light source useful for the cultivation of solanaceous plants is known in the art. Accordingly, the spectral composition of the non-UV light source useful in the context of the present invention comprises 0-40%B, 0-40%G, 50-100%R and 0-40%Fr, preferably 10-20%B, 2-10%G, 70-90%R and 15-15%Fr, more preferably 12-16%B, 3-7%G, 75-85%R and 11-15%Fr.
[0100] In one aspect, the spectral composition of the light sources comprised in the system of the present invention (i.e. the spectral composition of all the light sources which irradiate the cultivated solanaceous plants in the plant cultivations area) results in a phytochrome stationary state (PSS) value which is particularly useful in solanaceous plant cultivation. Such PSS value useful in solanaceous plant cultivation is known in the art. In one aspect, the spectrum of the sources comprised in the system of the present invention results in a phytochrome stationary state (PSS) value of 0.70 to 0.90, preferably of 0.82 to 0.90, more preferably of 0.82 to 0.88, most preferably of 0.83 to 0.87.
[0101] The agricultural system for cultivating solanaceous plants according to the present invention comprises means required for cultivating solanaceous plants. Such means required for cultivating solanaceous plants are capable of providing the cultivation conditions suitable for the cultivation of solanaceous plants and are very well known in the art. Accordingly, means required for cultivating solanaceous plants comprised in the agricultural system for cultivating solanaceous plants according to the present invention may comprise, but are not limited to, one or more of means for controlling temperature, means for controlling humidity, means for controlling carbon dioxide (CO2), means for controlling shading, means for controlling ventilation, means for controlling irrigation, and means for distribution of nutrients to the cultivated solanaceous plants. In one aspect, the system according to the present invention further comprises one or more means suitable for controlling temperature, humidity, carbon dioxide (CO2) content, shading, ventilation, irrigation, and nutrient distribution.
[0102] The agricultural system for cultivating solanaceous plants according to the present invention accordingly may comprise a subassembly suitable for controlling the temperature in the plant cultivation area to a temperature range that is particularly suitable for cultivating solanaceous plants. The temperature range suitable for cultivating solanaceous plants is well known in the art. In one aspect, the temperature range suitable for cultivating solanaceous plants comprises a temperature in the range of 14 to 30 °C, preferably in the range of 16 to 28 °C, more preferably in the range of 18 to 27 °C, even more preferably in the range of 20 to 26 °C, or most preferably in the range of 22 to 25 °C. Preferably, the night temperature in the plant cultivation space is lower than the day temperature in the plant cultivation space.
[0103] The agricultural system for cultivating solanaceous plants according to the present invention accordingly may comprise a subassembly suitable for controlling the relative humidity of the atmosphere in the plant cultivation area to a relative humidity range that is particularly suitable for cultivating solanaceous plants. The relative humidity range suitable for cultivating solanaceous plants is well known in the art. In one aspect, the relative humidity range suitable for cultivating solanaceous plants comprises a relative humidity of the atmosphere in the range of 40 to 95%, preferably in the range of 50 to 85%, or more preferably in the range of 60 to 80%. Preferably, the night relative humidity of the atmosphere in the plant cultivation space is higher than the day humidity relative humidity of the atmosphere in the plant cultivation space.
[0104] The agricultural system for cultivating solanaceous plants according to the present invention accordingly may comprise a subassembly suitable for controlling the average CO2 content of the atmosphere in the plant cultivation area to a CO2 content that is particularly suitable for cultivating solanaceous plants. The CO2 content that is particularly suitable for cultivating solanaceous plants is well known in the art. IN one aspect, the CO2 content that is particularly suitable for cultivating solanaceous plants comprises a CO2 content in the range of 300 to 2000 ppm, preferably in the range of 300-1200 ppm, more preferably in the range of 400 to 1100 ppm, even more preferably in the range of 500 to 1000 ppm, or most preferably in the range of 600 to 800 ppm. Preferably, the night CO2 content of the atmosphere in the plant cultivation area is lower than the day CO2 content of the atmosphere in the plant cultivation area.
[0105] The agricultural system for cultivating solanaceous plants according to the present invention accordingly may comprise a subassembly suitable for controlling the shading in the plant cultivation area. Means suitable for controlling the shading in a plant cultivation area are well known in the art including, but are not limited to, netting and roller blinds.
[0106] The agricultural system for cultivating solanaceous plants according to the present invention accordingly may comprise a subassembly suitable for controlling the ventilation in the plant cultivation area. Means suitable for controlling ventilation in a plant cultivation area are well known in the art including, but are not limited to, vents and ventilators.
[0107] The agricultural system for cultivating solanaceous plants according to the present invention accordingly may comprise a subassembly suitable for controlling the irrigation provided to the plants to maintain plant cultivation conditions that are particularly suitable for cultivating solanaceous plants. Means suitable for irrigation of plants in a plant cultivation area are well known in the art including, but are not limited to, sprinkling irrigation, misting irrigation and drip irrigation.
[0108] The agricultural system for cultivating solanaceous plants according to the present invention accordingly may comprise a subassembly suitable for controlling the nutrients provided to the plants to maintain plant cultivation conditions that are particularly suitable for cultivating solanaceous plants. Means suitable for controlling the nutrients provided to plants in a plant cultivation area are well known in the art including. The nutrient distribution system preferably is integrated in the irrigation system, most preferably integrated in the drip irrigation system.
[0109] The agricultural system for cultivating solanaceous plants according to the present invention may further comprise a control unit which controls one or more subassemblies comprised in the system of the present invention, wherein said one or more subassemblies comprise, but are not limited to: a means suitable for controlling temperature, a means suitable for humidity, a means suitable for carbon dioxide (CO2) content, a means suitable for shading, a means suitable for ventilation, a means suitable for irrigation, and a means suitable for nutrient distribution.
[0110] The agricultural system according to the present invention comprises a plant cultivation area, which accordingly comprises a plant cultivation substrate providing a suitable support for the root system of the plants cultivated in the plant cultivation space. Plant cultivation substrates suitable for the cultivation of solanaceous plants are well known in the art. In one aspect, the system according to the present invention comprises a plant cultivation substrate in the plant cultivation area, wherein said plant cultivation substrate preferably comprises one or more of soil, mineral wool, vermiculite and perlite.Method for cultivating solanaceous plants
[0111] The present invention further provides a method for cultivating plants that is characterized in that the solanaceous plants are cultivated in the system for cultivating solanaceous plants of the present invention. All technical features as described herein in the context of the agricultural system for cultivating solanaceous plants according to the present invention may also be applied in the method for cultivating solanaceous plants according to the present invention.
[0112] The present invention accordingly provides a method for cultivating solanaceous plants in a controlled plant cultivation environment deficient in solar UV radiation, said method comprises subjecting the plant to supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of 300 to 322 nm.
[0113] The method for cultivating solanaceous plants according to the present invention accordingly comprises subjecting the plant to supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of 300 to 322 nm. Preferably, the method for cultivating solanaceous plants according to the present invention accordingly comprises subjecting the plant to supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of 300 to 320 nm, 300 to 318 nm, 300 to 316 nm, 300 to 315 nm, 300 to 314 nm, 300 to 313 nm, 300 to 312 nm, 300 to 311 nm, 300 to 310 nm, 301 to 322 nm, 301 to 320 nm, 301 to 318 nm, 301 to 316 nm, 301 to 315 nm, 301 to 314 nm, 301 to 313 nm, 301 to 312 nm, 301 to 311 nm, 301 to 310 nm, 302 to 322 nm, 302 to 320 nm, 302 to 318 nm, 302 to 316 nm, 302 to 315 nm, 302 to 314 nm, 302 to 313 nm, 302 to 312 nm, 302 to 311 nm, 302 to 310 nm, 303 to 322 nm, 303 to 320 nm, 303 to 318 nm, 303 to 316 nm, 303 to 315 nm, 303 to 314 nm, 303 to 313 nm, 303 to 312 nm, 303 to 311 nm, 303 to 310 nm, 304 to 322 nm, 304 to 320nm, 304 to 318 nm, 304 to 316 nm, 304 to 315 nm, 304 to 314 nm, 304 to 313 nm, 304 to 312 nm, 304 to 311 nm, 304 to 310 nm, 305 to 322 nm, 305 to 320 nm, 305 to 318 nm, 305 to 316 nm, 305 to 315 nm, 305 to 314 nm, 305 to 313 nm, 305 to 312 nm, 305 to 311 nm, 305 to 310 nm, 306 to 322 nm, 306 to 320 nm, 306 to 318 nm, 306 to 316 nm, 306 to 315 nm, 306 to 314 nm, 306 to 313 nm, 306 to 312 nm, 306 to 311 nm or 306 to 310 nm. More preferably, the method for cultivating solanaceous plants according to the present invention accordingly comprises subjecting the plant to supplemental UV radiation emitted by an LIV-LED light source having a peak wavelength in the range of 305 to 315 nm.
[0114] Preferably, the method for cultivating solanaceous plants according to the present invention does not comprise subjecting the cultivated plants to supplemental UV emitted by a gas discharge UV light source. More preferably, the method for cultivating solanaceous plants does not comprise subjecting the cultivated plants to supplemental UV emitted by other supplemental UV light sources than the UV-LED light source according to the present invention.
[0115] The present invention accordingly provides a method for cultivating solanaceous plants. Preferably, the solanaceous plants cultivated in the method according to the present invention are potato plants, tobacco plants, pepper plants, eggplant or tomato plants. Most preferably, the solanaceous plants cultivated in the method according to the present invention are tomato plants.
[0116] In one aspect, the UV-LED light source that subjects the plant to supplemental UV radiation in the method for cultivating solanaceous plants according to the present invention is a narrow bandwidth UV-LED light source having a full width at half maximum (FWHM) emission spectrum of at most 30 nm. Preferably, the UV-LED light source that subjects the plant to supplemental UV radiation in the method for cultivating solanaceous plants according to the present invention is a narrow bandwidth UV-LED light source having a full width at half maximum (FWHM) emission spectrum of at most 29 nm, 28 nm, 27 nm, 26 nm, 25 nm, 24 nm, 23 nm, 22 nm, 21 nm, 20 nm, 19 nm, 18 nm, 17 nm, 16 nm, 15 nm, 14 nm, 13 nm, 12 nm, 11 nm or 10 nm.
[0117] Preferably, the method for cultivating solanaceous plants comprises subjecting the plant to supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of (i.e. a peak wavelength of) 300 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peakwavelength of 301 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 30nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 30 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 30 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 30 nm.
[0118] More preferably, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to supplemental UV radiation emitted by an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 27 nm, apeak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 27 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 27 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 27 nm.
[0119] Even more preferably, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to supplemental UV radiation emitted by an UV- LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peakwavelength of 301 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 25 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of atmost 25 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 25 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 25 nm.
[0120] Particularly preferably, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to supplemental UV radiation emitted by an UV- LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 22 nm, apeak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 22 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 22 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 22 nm.
[0121] More particularly preferably, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to supplemental UV radiation emitted by an UV- LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peakwavelength of 302 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 21 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of atmost 21 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 21 nm.
[0122] Even more particularly preferably, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to supplemental UV radiation emitted by an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM spectrum of at most 20 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 20 nm, apeak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 20 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 20 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 20 nm.
[0123] Most preferably, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to supplemental UV radiation emitted by an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm and a FWHM spectrum of at most 18 nm, a peak wavelength of 300 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 300 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 301 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 302 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 303 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 304 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 311 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 305 to 310 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 322 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 320 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 318 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 316 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 315 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 314 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 313 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 312 nm and a FWHM emission spectrum of at most 18 nm, a peak wavelength of 306 to 311 nm and a FWHM emission spectrum of at most 18 nm, or a peak wavelength of 306 to 310 nm and a FWHM emission spectrum of at most 18 nm.
[0124] In one aspect, the intensity of the UV radiation emitted by the LIV-LED light source that subjects the plant to supplemental UV radiation in the method for cultivating solanaceous plants according to the present invention is 0.1 to 2 pmol / m2 / s at plant height. Preferably, the intensity at plant height of the UV radiation emitted by the UV-LED light source that subjects the plant to supplemental UV radiation in the method for cultivating solanaceous plants according to the present invention is 0.1 to 2 pmol / m2 / s, 0.2 to 2 pmol / m2 / s, 0.3 to 2 pmol / m2 / s, 0.1 to 1.5 pmol / m2 / s, 0.2 to 1.5 pmol / m2 / s, 0.3 to 1.5 pmol / m2 / s, 0.1 to 1 pmol / m2 / s, 0.2 to 1 pmol / m2 / s or 0.3 to 1 pmol / m2 / s. Most preferably, the intensity at plant height of the UV radiation emitted by the UV- LED light source that subjects the plant to supplemental UV radiation in the method for cultivating solanaceous plants according to the present invention is 0.3 to 1.5 pmol / m2 / s.
[0125] Accordingly, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of (i.e. a peak wavelength of) 300 to 322 nm, a FWHM emission spectrum of at most (i.e. a FWHM spectrum of at most) 30 nm and an intensity at plant height of (i.e. intensity of) 0.1 to 2 pmol / m2 / s. Preferably, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to supplemental UV radiation emitted by an UV-LED light source having: a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.1 to 2 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.2 to 1 .5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.2 to 1 .5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 30 nm and anintensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.2 to 1 .5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.2 to 1 .5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.2 to 1 .5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.2 to 1 .5 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.2 to 1.5 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 30 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 27 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 25 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 22 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 322 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 320 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.3 to 1 pmol / m2 / s; a peak wavelength of 300 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.3 to 1 pmol / m2 / s; or a peak wavelength of 305 to 315 nm, a FWHM spectrum of at most 20 nm and an intensity of 0.3 to 1 pmol / m2 / s.
[0126] In the method for cultivating solanaceous plants according to the present the solanaceous plants the solanaceous plants are subjected to conditions suitable for the cultivation of solanaceous plants. Said suitable conditions for the cultivation of solanaceous plants comprises subjecting the solanaceous plants to an effective amount of non-UV light. Said non-UV light may originate from the sun (e.g. in a glass greenhouse) and / or from an artificial light source. An effective amount of non-UV light is characterized in that the cultivated plant is subjected to non-UV light having an intensity and spectral composition which is suitable for solanaceous plant cultivation. Such non-UV light having a spectral composition and intensity useful in solanaceous plant cultivation is known in the art.
[0127] In one aspect, the method for cultivating solanaceous plants according to the present invention further comprises subjecting the plant to supplemental non-UV light emitted by an artificial non-UV light source. Preferably, the artificial non-UV light source is a "non-UV-LED light source".
[0128] In one aspect, the supplemental non-UV light in the method according to the present invention comprises an intensity at plant height of 5-1000 pmol / m2 / s. Preferably, the supplemental non- UV light in the method according to the present invention comprises an intensity at plant height of 5-1000 pmol / m2 / s, 10-1000 pmol / m2 / s, 15-1000 pmol / m2 / s, 20-1000 pmol / m2 / s, 25-1000 pmol / m2 / s, 30-1000 pmol / m2 / s, 35-1000 pmol / m2 / s, 40-1000 pmol / m2 / s, 50-1000 pmol / m2 / s, 60- 1000 pmol / m2 / s, 70-1000 pmol / m2 / s, 80-1000 pmol / m2 / s, 90-1000 pmol / m2 / s, 100-1000 pmol / m2 / s, 5-500 pmol / m2 / s, 10-500 pmol / m2 / s, 15-500 pmol / m2 / s, 20-500 pmol / m2 / s, 25-500 pmol / m2 / s, 30-500 pmol / m2 / s, 35-500 pmol / m2 / s, 40-500 pmol / m2 / s, 50-500 pmol / m2 / s, 60-500 pmol / m2 / s, 70-500 pmol / m2 / s, 80-500 pmol / m2 / s, 90-500 pmol / m2 / s, 100-500 pmol / m2 / s, 5-250 pmol / m2 / s, 10-250 pmol / m2 / s, 15-250 pmol / m2 / s, 20-250 pmol / m2 / s, 25-250 pmol / m2 / s, 30-250 pmol / m2 / s, 35-250 pmol / m2 / s, 40-250 pmol / m2 / s, 50-250 pmol / m2 / s, 60-250 pmol / m2 / s, 70-250 pmol / m2 / s, 80-250 pmol / m2 / s, 90-250 pmol / m2 / s, 100-250 pmol / m2 / s, 5-200 pmol / m2 / s, 10-200 pmol / m2 / s, 15-200 pmol / m2 / s, 20-200 pmol / m2 / s, 25-200 pmol / m2 / s, 30-200 pmol / m2 / s, 35-200 pmol / m2 / s, 40-200 pmol / m2 / s, 50-200 pmol / m2 / s, 60-200 pmol / m2 / s, 70-200 pmol / m2 / s, 80-200 pmol / m2 / s, 90-200 pmol / m2 / s, 100-200 pmol / m2 / s, 5-180 pmol / m2 / s, 10-180 pmol / m2 / s, 15-180 pmol / m2 / s, 20-180 pmol / m2 / s, 25-180 pmol / m2 / s, 30-180 pmol / m2 / s, 35-180 pmol / m2 / s, 40-180 pmol / m2 / s, 50-180 pmol / m2 / s, 60-180 pmol / m2 / s, 70-180 pmol / m2 / s, 80-180 pmol / m2 / s, 90-180 pmol / m2 / s, 100-180 pmol / m2 / s, 5-170 pmol / m2 / s, 10-170 pmol / m2 / s, 15-170 pmol / m2 / s, 20-170 pmol / m2 / s, 25-170 pmol / m2 / s, 30-170 pmol / m2 / s, 35-170 pmol / m2 / s, 40-170 pmol / m2 / s, 50-170 pmol / m2 / s, 60-170 pmol / m2 / s, 70-170 pmol / m2 / s, 80-170 pmol / m2 / s, 90-170 pmol / m2 / s, 100-170 pmol / m2 / s, 5-160 pmol / m2 / s, 10-160 pmol / m2 / s, 15-160 pmol / m2 / s, 20-160 pmol / m2 / s, 25-160 pmol / m2 / s, 30-160 pmol / m2 / s, 35-160 pmol / m2 / s, 40-160 pmol / m2 / s, 50-160 pmol / m2 / s, 60-160 pmol / m2 / s, 70-160 pmol / m2 / s, 80-160 pmol / m2 / s, 90-160 pmol / m2 / s, 100-160 pmol / m2 / s, 5-150 pmol / m2 / s, 10-150 pmol / m2 / s, 15-150 pmol / m2 / s, 20-150 pmol / m2 / s, 25-150 pmol / m2 / s, 30-150 pmol / m2 / s, 35-150 pmol / m2 / s, 40-150 pmol / m2 / s, 50-150 pmol / m2 / s, 60-150 pmol / m2 / s, 70-150 pmol / m2 / s, 80-150 pmol / m2 / s, 90-150 pmol / m2 / s, or 100-150 pmol / m2 / s.
[0129] In one aspect, the supplemental non-UV light in the method according to the present invention comprises: a Blue light component (B) having a peak wavelength of more than 400 nm to at most 495 nm (preferably 435-485 nm), a Green light component (G) having a peak wavelength of more than 495 nm to at most 590 nm, (preferably 500-550 nm), a Red light component (R) having a peak wavelength of more than 590 nm to at most 700 nm (preferably 600-650 nm) and a Far-red light component (Fr) having a peak wavelength of more than 700 to at most 800 nm.The spectral composition of the non-UV light source useful for the cultivation of solanaceous plants is known in the art. Accordingly, the spectral composition of the non-UV light source useful in the context of the present invention comprises 0-40%B, 0-40%G, 50-100%R and 0-40%Fr, preferably 10-20%B, 2-10%G, 70-90%R and 15-15%Fr, more preferably 12-16%B, 3-7%G, 75- 85%R and 11-15%Fr.
[0130] In one aspect, the spectral composition of the light administered to the plants in the method of the present invention results in a phytochrome stationary state (PSS) value which is particularly useful in solanaceous plant cultivation. Such PSS value useful in solanaceous plant cultivation is known in the art. In one aspect, the spectral composition of the light administered to the solanaceous plants results in a phytochrome stationary state (PSS) value of 0.70 to 0.90, preferably of 0.82 to 0.90, more preferably of 0.82 to 0.88, most preferably of 0.83 to 0.87.
[0131] The method for cultivating solanaceous plants according to the present invention may further comprise a photoperiod, which refers to a period of the day in which the plants are subjected to light (“photoperiod”) in addition to a period wherein the plants are not subjected to light (“night period”). In one aspect, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to light for 10-20 hrs per day, preferably 12-18 hrs per day, more preferably 14-16 hrs per day. In one aspect, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to non-UV light for 10-20 hrs per day, preferably 12-18 hrs per day, more preferably 14-16 hrs per day. The method for cultivating solanaceous plants according to the present invention may further comprise an UV photoperiod, which in the context of the present invention refers to a period of the day in which the plants are subjected to UV light (“UV photoperiod”) in addition to a period wherein the plants are not subjected to UV light. In one aspect, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plant to supplemental UV radiation for 1-24 hrs per day, preferably 4-20 hrs per day, more preferably 6-18 hrs per day, even more preferably 8-16 hrs per day. In one aspect, the method for cultivating solanaceous plants according to the present invention comprises subjecting the plants to supplemental UV radiation for a shorter period per day than subjecting the plants to non-UV light. Preferably, the method for cultivating solanaceous plants according to the present invention comprises switching off the UV-LED light source when humans are present in the plant growth space, e.g. by using an automated system like a clock switch or a motion detector switch.
[0132] The suitable conditions for the cultivation of solanaceous plants comprised in the method for cultivating solanaceous plants according to the present invention may further comprise a temperature suitable for cultivation of solanaceous plants, a humidity suitable for cultivation of solanaceous plants, a carbon dioxide (CO2) content suitable for cultivation of solanaceous plants, irrigation suitable for cultivation of solanaceous plants and nutrient distribution suitable for cultivation of solanaceous plants. Temperature suitable for cultivation of solanaceous plants, humidity suitable for cultivation of solanaceous plants, carbon dioxide (CO2) content suitable for cultivation of solanaceous plants, irrigation suitable for cultivation of solanaceous plants and nutrient distribution suitable for cultivation of solanaceous plants are well known in the art. Suitable temperature ranges, suitable relative humidity ranges and suitable carbon dioxide (CO2) content ranges for solanaceous plant cultivation in the context of the method of the present invention are described herein in more detail in the context of the agricultural system of the present invention. Preferably, the method for cultivating solanaceous plants according to the present inventioncomprises cultivating the solanaceous plants under controlled conditions. In one aspect, the sol- anaceous plants in the method according to the present invention are cultivated under controlled conditions comprising one or more of: a temperature in the range of 14 to 30 °C; a relative humidity in the range of 40 to 95%; and a carbon dioxide content in the range of 300 to 2000 ppm. Preferably, the solanaceous plants in the method according to the present invention are cultivated under controlled conditions comprising one or more of: a temperature in the range of 16 to 28 °C; a relative humidity in the range of 50 to 85%; and a carbon dioxide content in the range of 300 to 1200 ppm.
[0133] In the context of the present invention, the solanaceous plants are cultivated on a plant cultivation substrate providing a suitable support for the root system of the solanaceous plants. The method according to the present invention accordingly comprises cultivation of solanaceous plants on a plant cultivation substrate, wherein said plant cultivation substrate preferably comprises one or more of soil, mineral wool, vermiculite and perlite. Accordingly, the solanaceous plants may be cultivated in soil (full soil or in pots), perlite bags, on an aerated nutrient solution, or on rockwool, or hydroponically (e.g., on sawdust or rockwool).
[0134] In one aspect, the present invention provides the use of supplemental UV radiation emitted by an LIV-LED light source having a peak wavelength in the range of 300 to 322 nm for the cultivation of solanaceous plants. Said use of supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of 300 to 322 nm for the cultivation of solanaceous plants is associated with numerous advantageous technical effects including, but not limited to, reducing intumescence injury of solanaceous plants cultivated in a plant cultivation area which is deficient in solar UV radiation.
[0135] The present invention accordingly provides the use of supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of 300 to 322 nm for reducing intumescence injury of solanaceous plants cultivated in a plant cultivation area which is deficient in solar UV radiation.
[0136] All technical features as described herein in the context of the herein described agricultural system for cultivating solanaceous plants and in the context of the method for cultivating solanaceous plants according to the present invention may also be advantageously applied in the uses according to the present invention.
[0137] The solanaceous plants used in the context of the present invention may be cultivated for the production and harvest of fruits. Said fruits may be particularly useful as food. Furthermore, said fruits may be useful for the seeds developed in the fruits. In the latter case, the solanaceous plants used in the context of the present invention may be cultivated for the production of seed. The method present invention may be useful for producing seeds of any solanaceous plant that is susceptible for intumescence injury when cultivated in a controlled plant cultivation environment deficient in solar UV radiation, wherein the seeds are produced by a seed production plant. The method and use for cultivating solanaceous plants according to the present invention accordingly is particularly useful for producing plant seeds in a controlled plant cultivation environment deficient in solar UV radiation. Accordingly, the present invention is particularly useful for producing seeds in a tunnel, in a glass green house or in a climate chamber. In one aspect, accordingly all seed production plants are of the same female parent line, preferably the same inbred femaleparent line. Likewise, it is understood that the male parent plant which was previously used for pollination of the female parent plant is of one genotype, e.g. of a male parent line, e.g. a male parent inbred line. In one aspect, the seed production plant in the method or use of the present invention is an inbred plant. In one aspect, the seed production plant in the method or use of the present invention is an F1 hybrid parental line plant. In one aspect, the seed production plant in the method or use of the present invention is an emasculated plant. In one aspect, the seed production plant in the method or use of the present invention is a male sterile plant. In one aspect, the plant seeds produced by the seed production plant are F1 hybrid plant seeds. Such F1 hybrid plants may have advantages such as improved uniformity, vitality and / or disease tolerance.EXAMPLESEXAMPLE 1
[0138] In Example 1 it is determined which wavelength of UV-LED light is most effective at suppressing intumescence in tomato. The experiment was done in a climate room with four cabins with 2 m2net area. The set points and timeline of this experiment are shown in Table 1.1 and 1.2. LED lighting was used, but no artificial sunlight. A control without any UV light was compared with treatments where additional UV light of different wavelengths was supplied during the light period (Table 1.3). Lighting is described in detail herein below.Table 1.1. Set points.Table 1.2 TimelineTable 1.3 Treatments. All treatments had LED lighting as well
[0139] All four climate cabins had LED lighting with a spectrum of 14%B 5%G 81 %R 13%Fr relative to PAR, and PSS 0.86 (See Fig. 1 for the spectrum). There was no artificial sunlight. Photoperiod was 15 hours per day and intensity was 100 pmol / m2 / s from 0 das and 150 pmol / m2 / s from 21 das at plant height (Table 1.2).
[0140] Custom-made UV LED lights were used (I llumicent, Breda, Netherlands). The 285 nm UV- C LEDs were type XBT-3535-UV (Luminus Devices, Inc. Sunnyvale, CA) and the 308 nm and 325 nm LEDs were respectively MODEL 308-FL-02-U05 and MODEL 325-FL-02-U05 (DOWA Electronics Materials Co., Ltd). The spectrum is shown in Figure 2. UV light had an intensity of 0.12 pmol / m2 / s at plant height in all three treatments. UV lights were on during the light period only.
[0141] Seedlings 10 days after the start of treatments showed heavy leaf damage due to intumescence in the control without UV (Fig. 3). Seedlings in the 285-nm UV treatment were severely damaged and stunted. However, this was not the result of intumescence but damage by UV radiation at this particular wavelength close to the UV-C band. Such short wavelength UV radiation is known to be very damaging to living tissues. The 285-nm treatment was not continued further.
[0142] When treated with 308-nm UV light the seedlings were healthy and big, without any visible intumescence at this stage. Seedlings in the 325-nm UV treatment displayed mild intumescence symptoms but also appeared much better than the untreated control (Fig. 3).
[0143] One week later at 24 das there was still a large contrast in leaf damage between UV-treated plants and the control without any UV light. Figure 4 shows the lower (abaxial) side of a representative fully elongated leaf from the top of the plant. By this time intumescence had also developed in the 308-nm treated plants, but the leaves were much bigger than the untreated control. The 325-nm treated plants had a smaller leaf size than 308-nm treated plants but were also larger and less damaged by intumescence than the control.
[0144] After another 10 days, at 34 das, plants were photographed, measured for length and fresh weight, and the experiment was ended. Plants in the 308-nm UV treatment were much larger than the untreated control, while 325-nm UV treated plants were intermediate between the two (Fig. 5-6). Control plants were stunted, the lower leaves were damaged, and leaves were strongly curled and pointed downwards.
[0145] The first flowering truss had appeared in plants treated with 308-nm UV light and 325-nm UV light but not in the control without any UV light. Hence, fruit production can be expected to be much delayed or failed completely in this variety under these conditions if no supplemental UV light is given.
[0146] Figure 7 compares a fully elongated leaf from the top of the plant in control and UV treatments. The 308-nm UV treated plant had a larger leaf than the untreated control or the 325-nm UV treated plant. The 325-nm treated leaf was paler than the other two.
[0147] The base of the stem showed clear symptoms of intumescence in the control without UV and the 325-nm UV treated plants, but not in the 308-nm treated plants (Fig. 8). The stem of the latter was also thicker than the other two and still had the cotyledons attached, while they had fallen off in the other two.
[0148] Plant height at final harvest was lowest for the control plants without any UV light, followed by 325-nm UV treated plants. 308-nm UV treated plants were much larger, double the height of the control (Fig. 9).
[0149] Plants from the 308-nm UV treatment that were left over after thinning 24 das were transferred to the 285-nm UV treatment. These plants were severely damaged and had reduced cultivation compared with 308-nm treated plants at final harvest 34 das. This shows again that 285- nm UV light, which is in the UV-B band, is very damaging to tomato cultivation and not suitable to prevent intumescence injury.
[0150] The difference in aboveground fresh weight at final harvest was even more striking (Fig. 10). Plants treated with 308-nm UV light had 7 times the weight of control plants and 325-nm treated plants had double the weight of untreated control plants. These data show that UV wavelength is a major factor determining resistance to intumescence in this LED light spectrum.
[0151] Tomato plants of the variety Competition were severely damaged by intumescence in an indoor cultivation system with LED lighting. Plants were stunted, cotyledons and lower leaves abscised, there were lesions on the stem and leaves, and leaf size was reduced. The results showed that a low intensity of supplemental UV light could reduce the damage by intumescence, but this depended strongly on UV wavelength. A peak UV wavelength of 308 nm was optimal. Plants treated with UV light of this wavelength were the largest and didn’t show any intumescence damage on the base of the stem, but there was damage on the lower side of the leaves. A slightly shorter peak wavelength of 285 nm was not suitable to protect plants against intumescence because UV light of this short wavelength was itself extremely damaging to seedlings. The longer wavelength of 325 nm reduced intumescence symptoms compared with a control that did not receive any UV light, but it was much less effective than UV light of 308 nm. In particular the aboveground fresh weight demonstrated this point: In the 308-nm UV treatment fresh weight was 7 times that of the control but in the 325-nm UV treatment it was only 2 times that of the control. Thus, the bandwidth of effective UV wavelengths that suppress intumescence is surprisingly nar- row.EXAMPLE 2
[0152] In Example 2 it was determined which is the optimal intensity is of 308-nm UV light to suppress intumescence in tomato. Example 1 showed that these plants were severely damaged by intumescence in a LED spectrum of 14%B 5%G 81 %R 13%Fr relative to PAR, and PSS 0.86, and that 0.12 pmol / m2 / s 308-nm UV light was much more effective in suppressing intumescence than either a shorter wavelength of 285 nm or a longer wavelength of 325 nm. The set points and timeline of this Example are shown in Table 2.1 and 2.2. LED lighting was used, but no artificial sunlight. A control without any UV light was compared with treatments where additional 308-nm UV light at different intensities was supplied during the light period (Table 2.3). Lighting is de- scribed in detail in Example 1 herein above.Table 2.1. Set points.Table 2.2 TimelineTable 2.3 Treatments. All treatments had LED lighting as well
[0153] Lighting was as in Example 1 except only 308-nm UV was used in different intensities. All four climate cabins had LED lighting with a spectrum of 14%B 5%G 81 %R 13%Fr relative to PAR, and PSS 0.86 (See Fig. 1 for the spectrum). There was no artificial sunlight.
[0154] Custom-made UV LED lights were used as in Example 1. The 308 nm LEDs were MODEL 308-FL-02-U05 (DOWA Electronics Materials Co., Ltd). The spectrum is shown in Figure 2. UV light intensities are in Table 2.3. UV lights were on during the light period only.
[0155] Plants in the control without UV again developed severe damage due to intumescence like in Example 1. They remained stunted with small downward curled leaves, the cotyledons and lower leaves abscised, and lesions were visible on the base of the stem (Fig. 11-14). By contrast, plants with 308-nm UV light at all intensities were much larger.
[0156] Plant height and fresh weight at final harvest 34 days after sowing significantly increased in all 308-nm UV treatments compared with control (Fig. 15, 16). Fresh weight increased by as much as 17-fold in the highest intensity 308-nm UV treatment compared with control.
[0157] Plant height increased when 308-nm UV intensity doubled from 0.12 to 0.24, but not with a further increase to 0.36 pmol / m2 / s. Above ground fresh weight increased with 308-nm UV intensity but between 0.24 and 0.36 pmol / m2 / s that was not statistically significant.
[0158] Compared with Example 1 , plants were smaller in control and 0.12 308nm UV, despite coming from the same seed batch and using the same cultivation conditions. However, the treatment effect was still clear and statistically significant.
[0159] Taken together, in these conditions, the intensity of 0.24 pmol / m2 / s 308-nm UV light was sufficient to suppress intumescence injury, increase leaf quality and size, plant height and fresh weight. It was more effective than 0.12 pmol / m2 / s.EXAMPLE 3
[0160] In Example 3 the effect was determined of a higher intensity of 308-nm UV light than the maximum intensity of 0.36 pmol / m2 / s in the previous experiment. Also, it is tested if the higher UV intensity causes any damage to the plants. The previous experiments showed that these plants were severely damaged by intumescence in a LED spectrum of 14%B 5%G 81%R 13%Fr relative to PAR, and PSS 0.86, and that 0.36 pmol / m2 / s 308-nm UV light strongly suppressed intumescence. The set points and timeline of this experiment are shown in Table 3.1 and 3.2. LED lightingwas used, but no artificial sunlight. A control without any UV light was compared with a treatment with additional 0.36 pmol / m2 / s 308-nm UV light and a higher intensity of 1.06 pmol / m2 / s 308-nm UV light supplied during the light period (Table 3.3). Lighting is described in detail herein below.Table 3.1. Set points.Table 3.2 TimelineTable 3.3 Treatments. All treatments had LED lighting as well
[0161] Lighting was as in Example 2 except for the UV light intensities. All three climate cabins hadLED lighting with a spectrum of 14%B 5%G 81 %R 13%Fr relative to PAR, and PSS 0.86 (See Fig. 1 for the spectrum). There was no artificial sunlight. Custom-made UV LED lights were used(I llumicent, Breda, Netherlands). The 308 nm LEDs were MODEL 308-FL-02-U05 (DOWA Electronics Materials Co., Ltd). The spectrum is shown in Figure 2 UV light intensities are in Table 3.3. UV lights were on during the light period only.
[0162] Plants in the control without UV again developed severe damage due to intumescence like in Examples 1 and 2. They remained stunted with small downward curled leaves, the cotyledons and lower leaves abscised, and lesions were visible on the base of the stem (see Figures. 17- 20). By contrast, plants with 308-nm UV light at both intensities were much larger, with bigger leaves, thicker stems without lesions and the cotyledons still attached.
[0163] At the medium intensity of 0.36 pmol / m2 / s 308-nm UV light intumescence was still visible but plants were larger, the leaves were larger and the stem was thicker than in the control without any UV light (see Figures. 18-20). At the highest intensity of 1.06 pmol / m2 / s 308-nm UV light plants developed normally and leaves appeared healthy and darker.
[0164] Plant height and fresh weight of leaves and stem at final harvest 31 days after sowing significantly increased in both 308-nm UV treatments compared with control (see Figures 21 and 22). Fresh weight of leaves and stem was significantly higher in 1.06 pmol / m2 / s 308-nm UV light than in 0.36 pmol / m2 / s 308-nm UV light, but plant height was similar.
[0165] Taken together, in these conditions, the intensity of 1.06 pmol / m2 / s 308-nm UV light improved plant quality further compared with 0.36 pmol / m2 / s 308-nm UV light by suppressing intumescence injury more. This resulted in almost no leaf damage and higher fresh weight of leaves and stem. No negative effects of the high intensity of 308-nm UV light on the plants were detected.
[0166] This Example 3 again showed that 308-nm UV light suppressed intumescence injury in the sensitive tomato variety Competition grown indoor. The first experiment already demonstrated that this wavelength of UV light was much more effective in suppressing intumescence than either the shorter wavelength or the longer wavelength in the UV range that was tested. The higher intensity of 1.06 pmol / m2 / s 308-nm UV light improved plant quality further compared with 0.36 pmol / m2 / s 308-nm UV light by suppressing intumescence injury more. This resulted in almost no leaf damage and higher fresh weight of leaves and stem. No negative effects of the high intensity of 308-nm UV light on the plants were detected.
Claims
CLAIMS1 . An agricultural system for cultivating solanaceous plants comprising a plant cultivation area which is deficient in solar UV radiation, wherein said system comprises an UV-LED light source having a peak wavelength in the range of 300 to 322 nm.
2. The system according to claim 1 , wherein the UV-LED light source is a narrow bandwidth UV-LED light source having a full width at half maximum (FWHM) emission spectrum of at most 30 nm.
3. The system according to claim 1 or 2, wherein the UV-LED light source during the UV photoperiod is capable of emitting UV radiation having an intensity of 0.1 to 2 pmol / m2 / s at plant height.
4. The system according to any one of the preceding claims, wherein said system further comprises an artificial non-UV light source, preferably a non-UV-LED light source.
5. The system according to claim 4, wherein the spectral composition of the light sources results in a phytochrome stationary state (PSS) value of 0.70 to 0.90.
6. The system according to any one of the preceding claims, wherein said system further comprises one or more means suitable for controlling temperature, humidity, carbon dioxide (CO2) content, shading, ventilation, irrigation, and nutrient distribution.
7. The system according to any one of the preceding claims, wherein plant cultivation substrate as comprised in the plant cultivation area comprises one or more of soil, mineral wool, vermiculite and perlite.
8. A method for cultivating solanaceous plants in a controlled plant cultivation environment deficient in solar UV radiation, said method comprises subjecting the plant to supplemental UV radiation emitted by an UV-LED light source having a peak wavelength in the range of 300 to 322 nm.
9. The method according to claim 8, wherein the UV-LED light source is a narrow bandwidth UV-LED light source having a full width at half maximum (FWHM) emission spectrum of at most 30 nm.
10. The method according to claim 8 or 9, wherein the intensity of the UV radiation emitted by the UV-LED light source is 0.1 to 2 pmol / m2 / s at plant height.
11. The method according to any one of the preceding claims, wherein said method further comprises subjecting the plant to supplemental non-UV light emitted by an artificial non-UV light source, preferably emitted by a non-UV-LED light source.
12. The method according to claim 11, wherein the spectral composition of the light administered to the solanaceous plants results in a phytochrome stationary state (PSS) value of 0.70 to 0.
90.
13. The method according to any one of the preceding claims, wherein the plant is subjected to supplemental UV radiation for 1 to 24 hrs per day.
14. The method according to any one of the preceding claims, wherein the plant is cultivated under controlled conditions comprising one or more of: a temperature in the range of 14 to 30 °C; a relative humidity in the range of 40 to 95%; and a carbon dioxide content in the range of 300 to 2000 ppm.
15. Use of supplemental UV radiation emitted by an UV-LED light source having a peak wave- length in the range of 300 to 322 nm for reducing intumescence injury of solanaceous plants cultivated in a plant cultivation area which is deficient in solar UV radiation.