Raspberry plant named ‘WSU 2188’

Abstract

This invention relates to a new and distinct variety of red raspberry plant (Rubus idaeus L.) named ‘WSU 2188’ from a controlled cross of two unnamed WSU raspberry selections that is different from its parents and other mid-season raspberries. ‘WSU 2188’ is primarily characterized by suitability for machine harvestability, adapted to the raspberry production area of the Pacific Northwest tolerance to root rot, mid-season production and large, conical fruit.

Description

STATEMENT REGARDING FEDERALLY-SPONSORED RESEARCH

[0001] This invention was made with government support under grant numbers 58-2072-9-027 and 58-5358-9-440 awarded by the United States Department of Agriculture through the Agricultural Research Service and grant number NI19HFPXXXXXG010 awarded by the United States Department of Agriculture through the National Institute of Food & Agriculture. The government has certain rights in the invention.

[0002] Latin name: This invention concerns a new and distinct cultivar of red raspberry plant with a botanical name of Rubus idaeus L.

[0003] Varietal denomination: The varietal denomination of the claimed raspberry is ‘WSU 2188.’FIELD OF THE INVENTION

[0004] This invention concerns a new and distinct cultivar of red raspberry plant with a botanical name of Rubus idaeus L. The intended market use for the fruit is as processed (frozen) or fresh.BACKGROUND OF THE INVENTION

[0005] Raspberries are an economically important crop in the United States and many cultivars of raspberry plant are known. New cultivars of raspberry are needed to provide growers with resources for emerging disease, climate, and market conditions that affect production.

[0006] The present invention relates to a new and distinct floricane-bearing (summer fruiting on second year canes) red raspberry variety designated as ‘WSU 2188’. The variety is botanically known as Rubus idaeus L. The new and distinct variety of red raspberry originated from a hand-pollinated cross of the ‘WSU 1638’ (unpatented) as the maternal parent and ‘WSU 1447’ (unpatented) as the paternal parent. ‘WSU 1638’ was selected from a cross made in 1999 and has good size and machine harvestability. ‘WSU 1447’ was selected from a cross made in 1996 and gives progeny that produce large, firm fruit suitable for machine harvesting.

[0007] The cross was made in Year 1 in Puyallup, WA. Seeds were germinated in the winter of Year 1-2 and the resulting seedlings planted in Year 2 with a cooperating raspberry grower in Burlington, WA. Selections were made in Year 4 using a raspberry machine harvester.

[0008] Varietal designation ‘WSU 2188’ was selected in Year 4 and asexually propagated in Puyallup, WA in tissue culture using meristem cultures from axillary buds of primocanes. This selection was planted in non-replicated plots with a cooperating grower in Year 5 and planted with a second grower in Year 8. The plantings were maintained by the growers using typical commercial growing practices and harvest methods. The plantings were subjectively evaluated weekly for adaptation to machine harvesting for two fruiting seasons. In all harvest seasons this selection machine harvested well and had large fruit size. This selection was tested for tolerance to root rot. Plants and fruit of this new variety have remained true to type through successive generations of asexual propagation. ‘WSU 2188’ is being named and released and is the subject of this invention.SUMMARY OF THE INVENTION

[0009] The following traits have been repeatedly observed and are determined to be unique characteristics of ‘WSU 2188’, which in combination distinguish this raspberry plant as a new and distinct variety:

[0010] 1. Large fruit size

[0011] 2. Conic fruit shape, with fruit longer than broad

[0012] 3. Tolerance to root rot

[0013] 4. Suitability for machine harvesting

[0014] 5. Long laterals and leaf stems that tend to droop

[0015] ‘WSU 2188’ is distinguished from its maternal parent, ‘WSU 1638’, by larger fruit and firmer fruit.

[0016] ‘WSU 2188’ is distinguished from its paternal parent, ‘WSU1447’, by larger fruit and softer fruit.

[0017] ‘WSU 2188’ is distinguished from ‘WSU 2166’ (U.S. Plant Pat. No. 30,980), by later fruiting, firmer fruit, larger fruit, darker fruit and lower tolerance to root rot.

[0018] ‘WSU 2188’ is distinguished from the most commonly grown red raspberry in the Pacific Northwest, ‘Meeker’ (unpatented), by larger fruit, firmer fruit, more drupelets per fruit, greater fruit length to width ratio and greater tolerance to root rot.

[0019] ‘WSU 2188’ is distinguished from ‘Willamette’ (unpatented) by larger fruit, greater fruit length to width ratio, later fruiting, lighter colored fruit and greater tolerance to root rot.BRIEF DESCRIPTION OF THE DRAWINGS

[0020] The accompanying photographs illustrate typical characteristics of the new variety ‘WSU 2188’ and are true to color as it is reasonably possible with color reproductions of this type. Color in the photographs may differ slightly from the color value cited in the detailed botanical description, which accurately describes the color of ‘WSU 2188’. When objects could be accurately measured with an 8 mm diameter measuring opening (leaves, fruit and some canes), color was measured with a Minolta CR-400 Colorimeter, which measures color in L*, a*, b* color coordinates. Calibration was performed using a standard white plate supplied by the manufacturer. These L*, a*, b* coordinates were converted and are presented in Munsell color notation. For all other color measurements, color was compared with Munsell (1977) color plates.

[0021] FIG. 1 is a picture of the growth habit of the plant and display of fruit on 16 June of Year 10, Lynden, WA.

[0022] FIG. 2 is a photograph of the primocane leaves of ‘WSU 2188’ on 22 July of Year 17, Lynden, WA.

[0023] FIG. 3 is a photograph of cane segments of ‘WSU 2188’, from left to right: basal segment of floricane, mid-section of floricane, basal segment of primocane, mid-section of primocane, and upper section of primocane, taken 4 August of Year 17, Lynden, WA.

[0024] FIG. 4 is a photograph of fruit and receptacle of ‘WSU 2188’ taken July 16 of Year 13 in Lynden, WA.

[0025] FIG. 5 is a photograph of machine harvested fruit of ‘WSU 2188’ taken 12 July of Year 10.

[0026] FIG. 6 is a photograph showing spring bud development of ‘WSU 2188’ relative to commonly grown cultivars taken 21 February Year 12, Puyallup, WA.DETAILED BOTANICAL DESCRIPTION

[0027] ‘WSU 2188’ has not been observed under all possible environmental conditions and as such the characteristics may vary in detail depending on weather conditions, day length, soil type and location. The description is based on observations taken in Year 11 (except as noted) in Puyallup, WA and with cooperating growers in Lynden, WA.

[0028] A comparison of fruit of ‘WSU 2188’ with cultivars commonly grown in Washington is given in Table 1. ‘WSU 2188’ had greater machine harvest yield than ‘Meeker’ and ‘Willamette’ and lower yield than ‘Cascade Harvest’. ‘WSU 2188’ had later midpoints of harvest than ‘Willamette’ and similar to ‘Meeker’. In the Puyallup planting, ‘WSU 2188’ had lower yield in Year 10 than ‘Cascade Harvest’ but greater than ‘Meeker’ and ‘Willamette’. In Year 11, yield of ‘WSU 2188’ was higher than all other cultivars. ‘WSU 2188’ had the largest fruit weight and firmest fruit of the genotypes tested. It had the lowest fruit rot in year 10 and second lowest in Year 11. ‘WSU 2188’ had midpoint of harvest a day earlier than ‘Meeker’ in Year 10 and four days later than ‘Meeker’ in Year 11.

[0029] TABLE 1Harvest data.‘WSU ‘Cascade2188’Harvest’‘Meeker’‘Willamette’Machine harvest, Lynden, WAzYield (tons / acre)Year 108.39.36.16.4Year 116.07.57.35.8Total14.316.813.412.2Midpoint of harvestYear 107 / 2 7 / 1 7 / 3 6 / 22Year 117 / 197 / 177 / 197 / 12Hand harvest, Puyallup, WAxYield (tons / acre)Year 108.111.06.87.8Year 117.27.06.56.3Total Yield15.318.013.314.1Fruit weight (g)Year 104.404.163.103.06Year 114.053.543.193.37Fruit firmness (g)Year 10102907474Year 11168107107122Fruit rot (%)Year 107.89.811.47.2Year 115.714.510.66.6Midpoint of harvestYear 106 / 276 / 236 / 286 / 19Year 117 / 137 / 117 / 9 7 / 5 zHarvest of non-replicated 8 plant plots.xHarvest of 3 replications of 3 plants.

[0030] Table 2 provides information on the plant and fruit characteristics of the new variety ‘WSU 2188’ compared with ‘Cascade Harvest’ and ‘Meeker’, varieties grown commercially in Washington.

[0031] TABLE 2Measurements of plant characteristics of ‘WSU 2188’ with two commercial varieties.‘WSU ‘Cascade 2188’Harvest’‘Meeker’Plant height (m)1.911.911.91Plant width (m)1.401.091.35Self-fertileyesyesyesFlower measurementsxDate of first bloom5 / 135 / 95 / 10Length of bloom (days)312831Number of petals555Length of petal (mm)7.076.096.01Width of petal (mm)3.743.504.13Color petal upper N9.5N9.5N9.5surface (Munsell)Color petal lower N9.5N9.5N9.5surface (Munsell)Petal shapeOblan-Oblan-Oblan-ceolateceolateceolatePetal marginEntireEntireEntirePetal TexturesmoothsmoothsmoothFlower diameter petal 20.217.917.8tip to petal tip at widest part of the flower (mm)Number of sepals555Color sepal upper 5GY 5 / 45GY 7 / 67.5GY 7 / 6surface (Munsell)Color sepal lower 5GY 5 / 45GY 7 / 85GY 7 / 6surface (Munsell)Length of sepal (mm)8.706.937.56Width of sepal (mm)6.255.555.40Flower diameter sepal 27.521.020.8tip to sepal tip at the widest part of the flower (mm)Number of stigma / flower677361Number of anthers / flower639265Flower fragrancenone none none notednotednotedPollen colorN 9.5N 9.5N 9.5Fruit MeasurementsFruit color (Munsell)5.3R 5.8R 5.1R 2.6 / 5.72.5 / 4.92.6 / 5.1Fruit weight (g)5.35.73.8Fruit length (mm)28.528.421.7Fruit width (mm)20.622.320.0Fruit length / width ratio1.391.271.09Pedicel length (mm)29.827.816.2Pedicel width (mm)1.01.01.0Receptacle length (mm)23.222.016.8Receptacle width (mm)8.18.99.0Drupelet length (mm)5.65.25.1Drupelet width (mm)4.14.53.7Drupelet weight (mg)49.751.133.1Number of seeds per fruit107112114Seed weight (mg)2.021.951.70PrimocanesNumber of primocanes / hill162618Length of primocane (m)2.122.192.55Diameter at ground (mm)9.511.51 1.9Diameter at 1.2 m (mm)7.99.88.8Number of nodes354350Internode length at 7.811.27.51.2 m (cm)Color primocane spines 2.5R 4 / 22.5R 4 / 22.5R 4 / 2(Munsell)Length spine (mm)1.92.31.5Length base of spine (mm)2.62.91.4Width base of spine (mm)1.11.40.9Color spine base (Munsell)2.5R 4 / 22.5R 4 / 25R 5 / 10Primary cane color 2.5GY 1.3GY 2.2GY (Munsell)7 / 66.5 / 3.96.6 / 4.0Secondary cane color 6YR 6.6R 7.9R (Munsell)6 / 53.8 / 4.83.9 / 4.5Primocane fruitingnononoPubescence on canesabsentabsentslightColor primocane leaves (Munsell)yTerminal LeafletUpper surface5.7GY 6.1GY n.d.3.7 / 4.73.5 / 3.8Lower surface4.6GY 5.0GY n.d.6.0 / 1.55.5 / 1.7Distal Lateral LeafletsUpper surface5.5GY 6.2GY n.d.4.0 / 4.23.4 / 3.7Lower surface4.6GY 5.1GY n.d.6.0 / 1.45.5 / 1.9Basal LeafletsUpper surface6.0GY 6.2GY n.d.3.6 / 3.63.3 / 3.8Lower surface4.7GY 5.9GY n.d.6.0 / 1.65.4 / 1.9Primocane leavesNumber of leaflets555Petiole length (mm)939989Rachis length (mm)395141Stipule (mm)6.78.311.3Leaflet marginsdoubly doubly doubly serratedserratedserratedLeaflet overlapoccasionallyoccasionallycommonlyoverlapoverlapoverlapTerminal leafletLength (mm)116104103Width (mm)566768Length / width2.11.51.5Leaflet tipcuspidatecuspidatecuspidateLeaflet baseobliquecordateobtuseDistal lateral leafletLength (mm)808378Width (mm)314139Length / width2.61.61.6Petiolule length (mm)01.61.6Leaflet tipcuspidatecuspidatecuspidateLeaflet baseobtuseobliqueobliqueBasal leafletLength (mm)969598Width (mm)525860Length / width1.81.61.6Petiolule length (mm)9.93.43.8Leaflet tipcuspidatecuspidatecuspidateLeaflet baseobtuseobliqueobliqueFloricanesPrimary cane color 2.3Y 1.3GY 2.2GY (Munsell)5.5 / 5.56.5 / 3.96.6 / 4.0Secondary cane 5.1YR 6.6R 7.9R color (Munsell)4.1 / 3.53.8 / 4.83.9 / 4.5Primocane fruitingnononoPubescence on canesabsentabsentslightNumber of floricanes / hill10.010.010.0Diameter at base (mm)10.59.811.4Diameter at 1.2 m (mm)7.97.611.1Internode length at 5.16.66.91.2 m (cm)Height (m)1.71.71.7Lateral length 1.2 m (cm)145146140Nodes / lateral12.211.312.7Number fruit / lateral19.021.015.0Number fruit / fruiting node3.53.72.1Color floricane leaveszTerminal LeafletUpper surface5.9GY 5.4GY n.d.3.4 / 3.83.5 / 3.4Lower surface4.4GY 5.5GY n.d.6.0 / 1.45.5 / 1.8Basal LeafletUpper surface3.9GY 6.4GY n.d.4.1 / 4.53.3 / 3.7Lower surface4.7GY 5.6GY n.d.5.7 / 1.65.6 / 1.8Floricane leavesLeaflet number333Petiole length (mm)424753Leaflet marginsdoublydoublydoublyserratedserratedserratedTerminal leafletLength (mm)998793Width (mm)678061Length / width1.51.11.5Petiolule (mm)28.030.224.5Basal leafletLength (mm)847876Width (mm)464843Length / width1.81.61.8Petiolule (mm)9.62.82.0xFlower measurements made in Year 13.yAverage color measurements for two leaflets of distal lateral and basal leaflets.zAverage color measurements for two leaflets of basal leaflets.

[0032] Machine harvested fruit was collected in Lynden, WA and analyzed in Years 7, 8 and 10. ‘WSU 2188’ had slightly lower soluble solids than ‘Meeker’ and higher titratable acidity and lower pH than ‘Meeker’ and ‘Willamette’, resulting in a slightly more tart flavor. Table 3 provides the comparative values for anthocyanin content, soluble solids, pH and titratable acidity of raspberry fruit for the ‘WSU 2188’, ‘Meeker’ and ‘Willamette’ varietals.

[0033] TABLE 3Anthocyanin content, soluble solids, pH and titratable acidity of raspberry fruit machine average of fruit harvested in years 2013, 2014 and 2017 at Lynden, WA. Based on fruit samples of approximately 300 g collected for each cultivar.WSUCascade2188PremierMeekerWillametteSoluble solids 9.17.59.48.4(deg. Brix)pH3.103.253.453.26Titratable acidityz 1.961.721.881.52(% citric acid)Total anthocyaninsy 82465795(mg C-3-G / 100 g FW)zAcidity expressed as % citric acid.yTotal anthocyanins determined spectrophotometrically from acidified ethanol extracts and expressed as cyanidin-3-glucoside.

[0034] Table 4 provides the root rot evaluation of ‘WSU 2188’, ‘Cascade Harvest’ and ‘Meeker,’ which were evaluated for susceptibility to root rot in plots at the Goss Farm, Puyallup, WA in plantings established in Year 8. The presence of Phytophthora fragariae var rubi ([W.F. Wilcox & J.M. Duncan] W.A. Man in't Veld,) in these root rot plots has been verified via PCR. Four plants of each clone were planted in this area. Plants were subjectively rated for vigor in the fall of each year from 0 to 5, with 0 being dead and 5 a healthy and vigorous plant. Year 11 was a very severe root rot test. Other raspberries in the same planting had a decrease in their vigor ratings by an average of over 40% from Year 10 to Year 11. In the fall of the fourth year, ‘WSU 2188’ plants had a rating of 3.5, while ‘Meeker’ averaged 2.5 and ‘Cascade Harvest’ averaged 1.0. Based on this trial, ‘WSU 2188’ demonstrates a relatively high root rot tolerance, although it is not absolutely unaffected.

[0035] TABLE 4Root rot evaluationx‘WSU ‘Cascade Date2188’Cascade Harvest’‘Meeker’Nov. 6, 20154.004.254.00Nov. 4, 20163.335.004.75Sep. 13, 20173.001.002.50xFour plants of each cultivar planted in Year 8. Subjectively rated 0-5. 0 = dead, 5 = healthy and vigorous.

[0036] ‘WSU 2188’ was planted in non-replicated plots with cooperating growers in Year 5 and Year 8. The plantings were maintained by growers using typical commercial methods and the fruit machine harvested. The plantings were subjectively evaluated for adaptation to machine harvesting weekly during the harvest season for two fruiting seasons. In all harvest seasons ‘WSU 2188’ machine harvested well and was productive with large fruit size. Based on these evaluations, ‘WSU 2188’ is suitable for machine harvesting. At most locations where ‘WSU 2188’ has been tested, no winter injury has been observed. In one location near Lynden, Washington, ‘WSU 2188’ showed a minimal amount of winter injury in one season. Drought and heat tolerance response are not known.

[0037] Table 5 provides fruit weight, firmness, and color measurements before and after storage for fruit of ‘WSU 2188’ and comparison cultivar Cascade Harvest. At harvest time the average fruit weight of WSU 2188 was 4.20 g before storage and 2.81 g after 14 days of storage. The average weight of ‘Cascade Harvest’ was 4.04 g before storage and 2.50 g after storage. The average firmness of ‘WSU 2188’ fruit before storage was 223.5 g and after storage was 68.5 g. The average firmness of ‘Cascade Harvest’ fruit before storage was 90.2 g and after storage was 27.0 g. ‘WSU 2188’ had larger, firmer and a darker red color (data not shown) than ‘Cascade Harvest’ both before storage and after storage.

[0038] TABLE 5Fruit weight and firmness before and after storage for hand-harvested fruit of two cultivars in plots Puyallup, WA.Fruit weight (g)Firmness (g)BeforeAfterBeforeAfterClonestoragezstoragestoragestorage‘WSU 2188’4.202.81223.568.5‘Cascade Harvest’4.042.50 90.227.0zValues represent means of 10 fruit per cultivar. Fruit were harvested 3 Jul. 2017, with data collected on individual fruit. Firmness was destructively measured on 10 fruit of each cultivar that were subsequently discarded. Fruits were stored at 4º C. for seven d, then brought to room temperature (~20° C.) for 4 h for after storage measurements.

Examples

Embodiment Construction

[0027]‘WSU 2188’ has not been observed under all possible environmental conditions and as such the characteristics may vary in detail depending on weather conditions, day length, soil type and location. The description is based on observations taken in Year 11 (except as noted) in Puyallup, WA and with cooperating growers in Lynden, WA.

[0028]A comparison of fruit of ‘WSU 2188’ with cultivars commonly grown in Washington is given in Table 1. ‘WSU 2188’ had greater machine harvest yield than ‘Meeker’ and ‘Willamette’ and lower yield than ‘Cascade Harvest’. ‘WSU 2188’ had later midpoints of harvest than ‘Willamette’ and similar to ‘Meeker’. In the Puyallup planting, ‘WSU 2188’ had lower yield in Year 10 than ‘Cascade Harvest’ but greater than ‘Meeker’ and ‘Willamette’. In Year 11, yield of ‘WSU 2188’ was higher than all other cultivars. ‘WSU 2188’ had the largest fruit weight and firmest fruit of the genotypes tested. It had the lowest fruit rot in year 10 and second lowest in Year 11....

STATEMENT REGARDING FEDERALLY-SPONSORED RESEARCH

[0001] This invention was made with government support under grant numbers 58-2072-9-027 and 58-5358-9-440 awarded by the United States Department of Agriculture through the Agricultural Research Service and grant number NI19HFPXXXXXG010 awarded by the United States Department of Agriculture through the National Institute of Food & Agriculture. The government has certain rights in the invention.

[0002] Latin name: This invention concerns a new and distinct cultivar of red raspberry plant with a botanical name of Rubus idaeus L.

[0003] Varietal denomination: The varietal denomination of the claimed raspberry is ‘WSU 2188.’FIELD OF THE INVENTION

[0004] This invention concerns a new and distinct cultivar of red raspberry plant with a botanical name of Rubus idaeus L. The intended market use for the fruit is as processed (frozen) or fresh.BACKGROUND OF THE INVENTION

[0005] Raspberries are an economically important crop in the United States and many cultivars of raspberry plant are known. New cultivars of raspberry are needed to provide growers with resources for emerging disease, climate, and market conditions that affect production.

[0006] The present invention relates to a new and distinct floricane-bearing (summer fruiting on second year canes) red raspberry variety designated as ‘WSU 2188’. The variety is botanically known as Rubus idaeus L. The new and distinct variety of red raspberry originated from a hand-pollinated cross of the ‘WSU 1638’ (unpatented) as the maternal parent and ‘WSU 1447’ (unpatented) as the paternal parent. ‘WSU 1638’ was selected from a cross made in 1999 and has good size and machine harvestability. ‘WSU 1447’ was selected from a cross made in 1996 and gives progeny that produce large, firm fruit suitable for machine harvesting.

[0007] The cross was made in Year 1 in Puyallup, WA. Seeds were germinated in the winter of Year 1-2 and the resulting seedlings planted in Year 2 with a cooperating raspberry grower in Burlington, WA. Selections were made in Year 4 using a raspberry machine harvester.

[0008] Varietal designation ‘WSU 2188’ was selected in Year 4 and asexually propagated in Puyallup, WA in tissue culture using meristem cultures from axillary buds of primocanes. This selection was planted in non-replicated plots with a cooperating grower in Year 5 and planted with a second grower in Year 8. The plantings were maintained by the growers using typical commercial growing practices and harvest methods. The plantings were subjectively evaluated weekly for adaptation to machine harvesting for two fruiting seasons. In all harvest seasons this selection machine harvested well and had large fruit size. This selection was tested for tolerance to root rot. Plants and fruit of this new variety have remained true to type through successive generations of asexual propagation. ‘WSU 2188’ is being named and released and is the subject of this invention.SUMMARY OF THE INVENTION

[0009] The following traits have been repeatedly observed and are determined to be unique characteristics of ‘WSU 2188’, which in combination distinguish this raspberry plant as a new and distinct variety:

[0010] 1. Large fruit size

[0011] 2. Conic fruit shape, with fruit longer than broad

[0012] 3. Tolerance to root rot

[0013] 4. Suitability for machine harvesting

[0014] 5. Long laterals and leaf stems that tend to droop

[0015] ‘WSU 2188’ is distinguished from its maternal parent, ‘WSU 1638’, by larger fruit and firmer fruit.

[0016] ‘WSU 2188’ is distinguished from its paternal parent, ‘WSU1447’, by larger fruit and softer fruit.

[0017] ‘WSU 2188’ is distinguished from ‘WSU 2166’ (U.S. Plant Pat. No. 30,980), by later fruiting, firmer fruit, larger fruit, darker fruit and lower tolerance to root rot.

[0018] ‘WSU 2188’ is distinguished from the most commonly grown red raspberry in the Pacific Northwest, ‘Meeker’ (unpatented), by larger fruit, firmer fruit, more drupelets per fruit, greater fruit length to width ratio and greater tolerance to root rot.

[0019] ‘WSU 2188’ is distinguished from ‘Willamette’ (unpatented) by larger fruit, greater fruit length to width ratio, later fruiting, lighter colored fruit and greater tolerance to root rot.BRIEF DESCRIPTION OF THE DRAWINGS

[0020] The accompanying photographs illustrate typical characteristics of the new variety ‘WSU 2188’ and are true to color as it is reasonably possible with color reproductions of this type. Color in the photographs may differ slightly from the color value cited in the detailed botanical description, which accurately describes the color of ‘WSU 2188’. When objects could be accurately measured with an 8 mm diameter measuring opening (leaves, fruit and some canes), color was measured with a Minolta CR-400 Colorimeter, which measures color in L*, a*, b* color coordinates. Calibration was performed using a standard white plate supplied by the manufacturer. These L*, a*, b* coordinates were converted and are presented in Munsell color notation. For all other color measurements, color was compared with Munsell (1977) color plates.

[0021] FIG. 1 is a picture of the growth habit of the plant and display of fruit on 16 June of Year 10, Lynden, WA.

[0022] FIG. 2 is a photograph of the primocane leaves of ‘WSU 2188’ on 22 July of Year 17, Lynden, WA.

[0023] FIG. 3 is a photograph of cane segments of ‘WSU 2188’, from left to right: basal segment of floricane, mid-section of floricane, basal segment of primocane, mid-section of primocane, and upper section of primocane, taken 4 August of Year 17, Lynden, WA.

[0024] FIG. 4 is a photograph of fruit and receptacle of ‘WSU 2188’ taken July 16 of Year 13 in Lynden, WA.

[0025] FIG. 5 is a photograph of machine harvested fruit of ‘WSU 2188’ taken 12 July of Year 10.

[0026] FIG. 6 is a photograph showing spring bud development of ‘WSU 2188’ relative to commonly grown cultivars taken 21 February Year 12, Puyallup, WA.DETAILED BOTANICAL DESCRIPTION

[0027] ‘WSU 2188’ has not been observed under all possible environmental conditions and as such the characteristics may vary in detail depending on weather conditions, day length, soil type and location. The description is based on observations taken in Year 11 (except as noted) in Puyallup, WA and with cooperating growers in Lynden, WA.

[0028] A comparison of fruit of ‘WSU 2188’ with cultivars commonly grown in Washington is given in Table 1. ‘WSU 2188’ had greater machine harvest yield than ‘Meeker’ and ‘Willamette’ and lower yield than ‘Cascade Harvest’. ‘WSU 2188’ had later midpoints of harvest than ‘Willamette’ and similar to ‘Meeker’. In the Puyallup planting, ‘WSU 2188’ had lower yield in Year 10 than ‘Cascade Harvest’ but greater than ‘Meeker’ and ‘Willamette’. In Year 11, yield of ‘WSU 2188’ was higher than all other cultivars. ‘WSU 2188’ had the largest fruit weight and firmest fruit of the genotypes tested. It had the lowest fruit rot in year 10 and second lowest in Year 11. ‘WSU 2188’ had midpoint of harvest a day earlier than ‘Meeker’ in Year 10 and four days later than ‘Meeker’ in Year 11.

[0029] TABLE 1Harvest data.‘WSU ‘Cascade2188’Harvest’‘Meeker’‘Willamette’Machine harvest, Lynden, WAzYield (tons / acre)Year 108.39.36.16.4Year 116.07.57.35.8Total14.316.813.412.2Midpoint of harvestYear 107 / 2 7 / 1 7 / 3 6 / 22Year 117 / 197 / 177 / 197 / 12Hand harvest, Puyallup, WAxYield (tons / acre)Year 108.111.06.87.8Year 117.27.06.56.3Total Yield15.318.013.314.1Fruit weight (g)Year 104.404.163.103.06Year 114.053.543.193.37Fruit firmness (g)Year 10102907474Year 11168107107122Fruit rot (%)Year 107.89.811.47.2Year 115.714.510.66.6Midpoint of harvestYear 106 / 276 / 236 / 286 / 19Year 117 / 137 / 117 / 9 7 / 5 zHarvest of non-replicated 8 plant plots.xHarvest of 3 replications of 3 plants.

[0030] Table 2 provides information on the plant and fruit characteristics of the new variety ‘WSU 2188’ compared with ‘Cascade Harvest’ and ‘Meeker’, varieties grown commercially in Washington.

[0031] TABLE 2Measurements of plant characteristics of ‘WSU 2188’ with two commercial varieties.‘WSU ‘Cascade 2188’Harvest’‘Meeker’Plant height (m)1.911.911.91Plant width (m)1.401.091.35Self-fertileyesyesyesFlower measurementsxDate of first bloom5 / 135 / 95 / 10Length of bloom (days)312831Number of petals555Length of petal (mm)7.076.096.01Width of petal (mm)3.743.504.13Color petal upper N9.5N9.5N9.5surface (Munsell)Color petal lower N9.5N9.5N9.5surface (Munsell)Petal shapeOblan-Oblan-Oblan-ceolateceolateceolatePetal marginEntireEntireEntirePetal TexturesmoothsmoothsmoothFlower diameter petal 20.217.917.8tip to petal tip at widest part of the flower (mm)Number of sepals555Color sepal upper 5GY 5 / 45GY 7 / 67.5GY 7 / 6surface (Munsell)Color sepal lower 5GY 5 / 45GY 7 / 85GY 7 / 6surface (Munsell)Length of sepal (mm)8.706.937.56Width of sepal (mm)6.255.555.40Flower diameter sepal 27.521.020.8tip to sepal tip at the widest part of the flower (mm)Number of stigma / flower677361Number of anthers / flower639265Flower fragrancenone none none notednotednotedPollen colorN 9.5N 9.5N 9.5Fruit MeasurementsFruit color (Munsell)5.3R 5.8R 5.1R 2.6 / 5.72.5 / 4.92.6 / 5.1Fruit weight (g)5.35.73.8Fruit length (mm)28.528.421.7Fruit width (mm)20.622.320.0Fruit length / width ratio1.391.271.09Pedicel length (mm)29.827.816.2Pedicel width (mm)1.01.01.0Receptacle length (mm)23.222.016.8Receptacle width (mm)8.18.99.0Drupelet length (mm)5.65.25.1Drupelet width (mm)4.14.53.7Drupelet weight (mg)49.751.133.1Number of seeds per fruit107112114Seed weight (mg)2.021.951.70PrimocanesNumber of primocanes / hill162618Length of primocane (m)2.122.192.55Diameter at ground (mm)9.511.51 1.9Diameter at 1.2 m (mm)7.99.88.8Number of nodes354350Internode length at 7.811.27.51.2 m (cm)Color primocane spines 2.5R 4 / 22.5R 4 / 22.5R 4 / 2(Munsell)Length spine (mm)1.92.31.5Length base of spine (mm)2.62.91.4Width base of spine (mm)1.11.40.9Color spine base (Munsell)2.5R 4 / 22.5R 4 / 25R 5 / 10Primary cane color 2.5GY 1.3GY 2.2GY (Munsell)7 / 66.5 / 3.96.6 / 4.0Secondary cane color 6YR 6.6R 7.9R (Munsell)6 / 53.8 / 4.83.9 / 4.5Primocane fruitingnononoPubescence on canesabsentabsentslightColor primocane leaves (Munsell)yTerminal LeafletUpper surface5.7GY 6.1GY n.d.3.7 / 4.73.5 / 3.8Lower surface4.6GY 5.0GY n.d.6.0 / 1.55.5 / 1.7Distal Lateral LeafletsUpper surface5.5GY 6.2GY n.d.4.0 / 4.23.4 / 3.7Lower surface4.6GY 5.1GY n.d.6.0 / 1.45.5 / 1.9Basal LeafletsUpper surface6.0GY 6.2GY n.d.3.6 / 3.63.3 / 3.8Lower surface4.7GY 5.9GY n.d.6.0 / 1.65.4 / 1.9Primocane leavesNumber of leaflets555Petiole length (mm)939989Rachis length (mm)395141Stipule (mm)6.78.311.3Leaflet marginsdoubly doubly doubly serratedserratedserratedLeaflet overlapoccasionallyoccasionallycommonlyoverlapoverlapoverlapTerminal leafletLength (mm)116104103Width (mm)566768Length / width2.11.51.5Leaflet tipcuspidatecuspidatecuspidateLeaflet baseobliquecordateobtuseDistal lateral leafletLength (mm)808378Width (mm)314139Length / width2.61.61.6Petiolule length (mm)01.61.6Leaflet tipcuspidatecuspidatecuspidateLeaflet baseobtuseobliqueobliqueBasal leafletLength (mm)969598Width (mm)525860Length / width1.81.61.6Petiolule length (mm)9.93.43.8Leaflet tipcuspidatecuspidatecuspidateLeaflet baseobtuseobliqueobliqueFloricanesPrimary cane color 2.3Y 1.3GY 2.2GY (Munsell)5.5 / 5.56.5 / 3.96.6 / 4.0Secondary cane 5.1YR 6.6R 7.9R color (Munsell)4.1 / 3.53.8 / 4.83.9 / 4.5Primocane fruitingnononoPubescence on canesabsentabsentslightNumber of floricanes / hill10.010.010.0Diameter at base (mm)10.59.811.4Diameter at 1.2 m (mm)7.97.611.1Internode length at 5.16.66.91.2 m (cm)Height (m)1.71.71.7Lateral length 1.2 m (cm)145146140Nodes / lateral12.211.312.7Number fruit / lateral19.021.015.0Number fruit / fruiting node3.53.72.1Color floricane leaveszTerminal LeafletUpper surface5.9GY 5.4GY n.d.3.4 / 3.83.5 / 3.4Lower surface4.4GY 5.5GY n.d.6.0 / 1.45.5 / 1.8Basal LeafletUpper surface3.9GY 6.4GY n.d.4.1 / 4.53.3 / 3.7Lower surface4.7GY 5.6GY n.d.5.7 / 1.65.6 / 1.8Floricane leavesLeaflet number333Petiole length (mm)424753Leaflet marginsdoublydoublydoublyserratedserratedserratedTerminal leafletLength (mm)998793Width (mm)678061Length / width1.51.11.5Petiolule (mm)28.030.224.5Basal leafletLength (mm)847876Width (mm)464843Length / width1.81.61.8Petiolule (mm)9.62.82.0xFlower measurements made in Year 13.yAverage color measurements for two leaflets of distal lateral and basal leaflets.zAverage color measurements for two leaflets of basal leaflets.

[0032] Machine harvested fruit was collected in Lynden, WA and analyzed in Years 7, 8 and 10. ‘WSU 2188’ had slightly lower soluble solids than ‘Meeker’ and higher titratable acidity and lower pH than ‘Meeker’ and ‘Willamette’, resulting in a slightly more tart flavor. Table 3 provides the comparative values for anthocyanin content, soluble solids, pH and titratable acidity of raspberry fruit for the ‘WSU 2188’, ‘Meeker’ and ‘Willamette’ varietals.

[0033] TABLE 3Anthocyanin content, soluble solids, pH and titratable acidity of raspberry fruit machine average of fruit harvested in years 2013, 2014 and 2017 at Lynden, WA. Based on fruit samples of approximately 300 g collected for each cultivar.WSUCascade2188PremierMeekerWillametteSoluble solids 9.17.59.48.4(deg. Brix)pH3.103.253.453.26Titratable acidityz 1.961.721.881.52(% citric acid)Total anthocyaninsy 82465795(mg C-3-G / 100 g FW)zAcidity expressed as % citric acid.yTotal anthocyanins determined spectrophotometrically from acidified ethanol extracts and expressed as cyanidin-3-glucoside.

[0034] Table 4 provides the root rot evaluation of ‘WSU 2188’, ‘Cascade Harvest’ and ‘Meeker,’ which were evaluated for susceptibility to root rot in plots at the Goss Farm, Puyallup, WA in plantings established in Year 8. The presence of Phytophthora fragariae var rubi ([W.F. Wilcox & J.M. Duncan] W.A. Man in't Veld,) in these root rot plots has been verified via PCR. Four plants of each clone were planted in this area. Plants were subjectively rated for vigor in the fall of each year from 0 to 5, with 0 being dead and 5 a healthy and vigorous plant. Year 11 was a very severe root rot test. Other raspberries in the same planting had a decrease in their vigor ratings by an average of over 40% from Year 10 to Year 11. In the fall of the fourth year, ‘WSU 2188’ plants had a rating of 3.5, while ‘Meeker’ averaged 2.5 and ‘Cascade Harvest’ averaged 1.0. Based on this trial, ‘WSU 2188’ demonstrates a relatively high root rot tolerance, although it is not absolutely unaffected.

[0035] TABLE 4Root rot evaluationx‘WSU ‘Cascade Date2188’Cascade Harvest’‘Meeker’Nov. 6, 20154.004.254.00Nov. 4, 20163.335.004.75Sep. 13, 20173.001.002.50xFour plants of each cultivar planted in Year 8. Subjectively rated 0-5. 0 = dead, 5 = healthy and vigorous.

[0036] ‘WSU 2188’ was planted in non-replicated plots with cooperating growers in Year 5 and Year 8. The plantings were maintained by growers using typical commercial methods and the fruit machine harvested. The plantings were subjectively evaluated for adaptation to machine harvesting weekly during the harvest season for two fruiting seasons. In all harvest seasons ‘WSU 2188’ machine harvested well and was productive with large fruit size. Based on these evaluations, ‘WSU 2188’ is suitable for machine harvesting. At most locations where ‘WSU 2188’ has been tested, no winter injury has been observed. In one location near Lynden, Washington, ‘WSU 2188’ showed a minimal amount of winter injury in one season. Drought and heat tolerance response are not known.

[0037] Table 5 provides fruit weight, firmness, and color measurements before and after storage for fruit of ‘WSU 2188’ and comparison cultivar Cascade Harvest. At harvest time the average fruit weight of WSU 2188 was 4.20 g before storage and 2.81 g after 14 days of storage. The average weight of ‘Cascade Harvest’ was 4.04 g before storage and 2.50 g after storage. The average firmness of ‘WSU 2188’ fruit before storage was 223.5 g and after storage was 68.5 g. The average firmness of ‘Cascade Harvest’ fruit before storage was 90.2 g and after storage was 27.0 g. ‘WSU 2188’ had larger, firmer and a darker red color (data not shown) than ‘Cascade Harvest’ both before storage and after storage.

[0038] TABLE 5Fruit weight and firmness before and after storage for hand-harvested fruit of two cultivars in plots Puyallup, WA.Fruit weight (g)Firmness (g)BeforeAfterBeforeAfterClonestoragezstoragestoragestorage‘WSU 2188’4.202.81223.568.5‘Cascade Harvest’4.042.50 90.227.0zValues represent means of 10 fruit per cultivar. Fruit were harvested 3 Jul. 2017, with data collected on individual fruit. Firmness was destructively measured on 10 fruit of each cultivar that were subsequently discarded. Fruits were stored at 4º C. for seven d, then brought to room temperature (~20° C.) for 4 h for after storage measurements.

Examples

Embodiment Construction

[0027]‘WSU 2188’ has not been observed under all possible environmental conditions and as such the characteristics may vary in detail depending on weather conditions, day length, soil type and location. The description is based on observations taken in Year 11 (except as noted) in Puyallup, WA and with cooperating growers in Lynden, WA.

[0028]A comparison of fruit of ‘WSU 2188’ with cultivars commonly grown in Washington is given in Table 1. ‘WSU 2188’ had greater machine harvest yield than ‘Meeker’ and ‘Willamette’ and lower yield than ‘Cascade Harvest’. ‘WSU 2188’ had later midpoints of harvest than ‘Willamette’ and similar to ‘Meeker’. In the Puyallup planting, ‘WSU 2188’ had lower yield in Year 10 than ‘Cascade Harvest’ but greater than ‘Meeker’ and ‘Willamette’. In Year 11, yield of ‘WSU 2188’ was higher than all other cultivars. ‘WSU 2188’ had the largest fruit weight and firmest fruit of the genotypes tested. It had the lowest fruit rot in year 10 and second lowest in Year 11....

Claims

1. A new and distinct variety of raspberry plant, designated ‘WSU2188’, substantially as illustrated and described herein.

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