31 results about "Glandularia" patented technology

Provided is a PagHB7a gene related to the salt tolerance and drought tolerance of poplar. The gene is used to create new germplasms of salt-tolerant and drought-tolerant 84K poplar (Populus alba×Populus glandulosa). Being a candidate gene related to the salt tolerance and drought resistance of poplar, the PagHB7a gene provides an important gene resource for breeding new varieties of salt-tolerant and drought-resistant poplar, and can be directly used for molecular-assisted breeding of stress-resistant poplar. In addition, a primer pair for amplifying the candidate gene and a transgenic modification method for poplar are provided, which can evaluate the stress resistance of poplar on the molecular level; on this basis, varieties with remarkable stress resistance can be accurately and efficiently screened, thus effectively shortening the breeding period of poplar.

The application provides a method for observing glandular hairs of Ainsliaea sinensis by using a scanning electron microscope. The method comprises the following steps: (1) selecting a region to be measured in a leaf blade, cutting a sample, and immersing the sample in an aldehyde fixing solution for fixation and immersion; (2) performing gradient dehydration by using an ethanol solution, performing replacement by using isopentyl acetate, and then performing drying by using a critical point drying instrument; (3) removing non-glandular hairs by using adhesive tape; and (4) then spraying gold and observing under a scanning electron microscope. By using the method, the observation and distribution density statistics of the glandular hairs of Ainsliaea sinensis can be simply, efficiently and accurately completed. By using the method, the problem of uneven leaf blade is well solved, and the problem of non-glandular hairs shielding and interfering with the visual field is completely solved.

The invention discloses an LjCYCD2 gene and application thereof in regulation and control of plant leaf glandular hair density, and belongs to the technical field of gene engineering. The amino acid sequence coded by the LjCYCD2 gene is as shown in SEQ ID NO: 2, and the nucleotide sequence of the LjCYCD2 gene is as shown in SEQ ID NO: 1. The gene is expressed in the honeysuckle leaf, the expression quantity is gradually reduced along with the maturity of the leaf, and after the LjCYCD2 gene of the honeysuckle leaf is silenced by VIGS, the gland hair density of the honeysuckle leaf is obviously reduced. The honeysuckle LjCYCD2 gene is used for constructing a plant overexpression vector and transforming tobacco, and the number of glandular hairs of leaves of the obtained transgenic tobacco is obviously increased. It can be seen that the honeysuckle LjCYCD2 gene is related to plant leaf glandular hair development and has important application value in regulation and control of plant leaf surface glandular hair density.

The invention relates to an NtWUN gene as well as an expression product, an expression vector and application of the NtWUN gene in regulation and control of plant glandular hair development, and aims to solve the technical problems that the development mechanism of different types of epidermal hairs in plants (such as tobacco) is unknown and effective molecular regulation and control measures and utilization ways are lacked at present. According to the invention, an NtWUN gene for negatively regulating epidermal hair development is identified, a gene editing vector and an overexpression vector are constructed, and a gene editing homozygous strain and an overexpression homozygous strain are obtained through screening; tissue expression, MeJA response mode analysis, subcellular localization, plant phenotypic characteristic identification and the like show that the NtWUN can negatively regulate the occurrence of epidermal hair and influence the change of the total density of the epidermal hair, and the NtWUN has important application value in regulating the development process of the plant epidermal hair.

The present invention discloses a method and apparatus for electrostatic separation of trichomes from plant biomass, addressing the limitations of conventional separation techniques. The method relies on differences in electrical properties of trichomes and plant material, utilizing an external electric field. A multi-component electrostatic separation assembly may prepare and separate one or more samples, thereby incorporating a dispensing component, a pipeline component, and a separation chamber comprising at least one electrode assembly. Moreover, the present invention may also provide for electrode assemblies that feature self-cleaning mechanisms.

The present invention relates to polynucleotide comprising a lncRNA gene according to SEQ ID No: 2, or according to a sequence having at least 50% sequence identity to SEQ ID No: 2, and an AP2c gene according to SEQ ID No: 3, or according to a sequence having at least 50% sequence identity to SEQ ID No: 3, which polynucleotide confers an increased density of type IV trichomes. The invention further relates to a plant comprising said polynucleotide and a SlASAT4 gene according to SEQ ID No. 21, or comprising a homologous sequence having at least 99.6% sequence identity to SEQ ID No. 21 wherein the presence of the polynucleotide leads to increased density of type IV trichomes and wherein the presence of said SlASAT4 gene leads to an increased ratio of S4:17 / S3:15 acyl sugars produced in glandular trichomes present on said plant when compared to a control plant lacking the polynucleotide and the SlASAT4 gene. Such a plant also was found to be resistant to Bemisia tabaci. The invention also discloses markers for the identification of the polynucleotide and the SlASAT4 gene and their use for the identification and development of Solanum lycopersicum plants. Also methods for identifying, producing and selecting of plants and seed are disclosed.

The invention discloses a plant glandular hair generation regulation gene NtMIXTA gene as well as an encoding protein, a biological material and application thereof, and aims to solve the technical problem of relatively low efficiency in the aspect of improving the density of plant glandular hair in the traditional breeding technology. According to the application, the NtMIXTA gene is cloned from tobacco, a knockout strain and an overexpression strain of the NtMIXTA gene are respectively constructed, and after the NtMIXTA gene is knocked out, the leaf surface glandular hair density of the plant is reduced, and the leaf surface glandular hair density of the overexpression strain is remarkably increased, which indicates that the NtMIXTA gene positively regulates the generation and development of the tobacco glandular hair. The application has important significance for researching a molecular regulatory network of plant glandular hair generation.

Glandular trichomes are epidermal outgrowths in plants that are the site of metabolic compound synthesis and storage. Modulating the secretion of metabolite compounds in tobacco and tea leaves can alter the taste profile in products made from these plants. The present disclosure relates to methods for modulating secretion from glandular trichomes and compositions comprising altered trichome secretion of metabolites.

The invention relates to a preparation method for simultaneously observing and counting glandular hairs, non-glandular hairs and stomata of mugwort leaves, and can effectively solve the problem that the distribution of glandular hairs, non-glandular hairs and stomata on the lower surface of mugwort leaves cannot be simultaneously observed. The technical solution is as follows: firstly, fresh mugwort leaves are taken, small pieces of leaves are cut from leaves with a knife, the leaves are placed in methanol for immersion and fixation, the leaves are rinsed with distilled water and then transferred to a sodium hypochlorite solution for bleaching until colorless, the leaves are rinsed with distilled water and then transferred to a tissue transparent agent for transparency, the materials are taken out and dyed with a safranin solution, the slides are sealed with water, and then observed with an optical microscope, the non-glandular hair handles and glandular hairs are dyed red, the distribution of glandular hairs, non-glandular hairs and stomata on the lower surface can be clearly observed and the counting is completed. The method of the invention has the characteristics of simple operation, low cost, small workload, high success rate and clear observation, and has low technical requirements for operators, and is an innovation in the preparation method.

The invention relates to the technical field of image recognition, and discloses a tobacco epidermis glandular hair measurement method based on a neural network, which comprises the following steps: a tobacco leaf processing step: taking a tobacco leaf plant which grows for 45-50 days and is completely unfolded, sampling is random sampling, and cutting is carried out along a leaf vein to obtain a sample; decoloring: putting the tobacco sample into a potassium hydroxide solution, and standing for a period of time to remove chlorophyll; a dyeing step: putting the decolored sample into rhodamine B dye liquor, and standing for a period of time to dye glandular hair; and a shooting step: shooting a sample image through a stereomicroscope to obtain a glandular hair image with a standard visual field size. By introducing the neural network technology, high-precision and full-process automatic operation of glandular hair identification and counting is realized, personal errors and workload of experimenters are greatly reduced, and by means of the deep learning technology, the system can continuously optimize the identification capability according to increase of the number and diversity of samples, so that the accuracy of glandular hair identification and counting is improved. The detection requirements of different types and forms of glandular hairs are met.

Glandular trichomes are epidermal outgrowths in plants that are the site of metabolic compound synthesis and storage. Modulating the secretion of metabolite compounds in tobacco and tea leaves can alter the taste profile in products made from these plants. The present disclosure relates to methods for modulating secretion from glandular trichomes and compositions comprising altered trichome secretion of metabolites.

The invention discloses an NtMIXTA-like gene, an encoding protein thereof, a biological material and application of the NtMIXTA-like gene in regulation and control of plant glandular trichome generation, and aims to solve the technical problem that an existing glandular trichome regulation and control gene is single in function. According to the application, the NtMIXTA-like gene is cloned from tobacco, a knockout plant and an overexpression plant of the NtMIXTA-like gene are respectively constructed, and the analysis on the leaf glandular hair of the NtMIXTA-like gene shows that the glandular hair density is obviously increased after the overexpression of the NtMIXTA-like gene, which indicates that the NtMIXTA-like gene can positively regulate and control the generation / development of the tobacco glandular hair. The application has important significance for researching a molecular regulatory network of plant glandular hair generation.

The invention provides a rapid cannabinoid content analysis method based on glandular hair density, and belongs to the technical field of content detection. Comprising the following steps: collecting high-definition glandular hair pictures of industrial hemp flowers and leaves, and calculating glandular hair density and cannabinoid content data in the industrial hemp flowers and leaves; carrying out correlation analysis by utilizing the two kinds of data to obtain a correlation model of the cannabinoid content and the glandular hair density in the industrial cannabis sativa flowers and leaves; calculating the glandular hair density of the industrial cannabis sativa floral leaf to be detected through the high-definition glandular hair picture, and substituting the glandular hair density into the correlation model of the cannabis sativa terpene content and the glandular hair density to quickly obtain the cannabis sativa terpene content in the industrial cannabis sativa floral leaf to be detected. The detection method is rapid, accurate and simple to operate, does not need any expensive precision equipment, only needs to shoot several high-definition glandular hair pictures, is suitable for real-time data detection of cannabinoid on site in field planting, is short in time from sample picture acquisition to data acquisition, and greatly improves the cannabinoid detection efficiency.

The present invention relates to a method, device and storage medium for identifying the leaf-fuzz ratio grade of moxibustion materials, which are applied to the field of traditional Chinese medicine technology, and include: obtaining the cellulose content of moxa floss and the length of non-glandular hairs of moxa floss. Through pre-experiment calculation, it can be known that the length and cellulose content of moxa floss have the highest contribution rate to various combustion indexes. That is to say, the heat release performance of moxa floss combustion is closely related to the length of non-glandular hairs and the cellulose content of moxa floss. And according to the moxa floss with known leaf-fuzz ratio grades, the length of non-glandular hairs and the cellulose content of different leaf-fuzz ratios can be known, that is, the standard value range of the length of preset non-glandular hairs and the standard value range of cellulose in this application. According to which leaf-fuzz ratio grade's standard value range the length of non-glandular hairs and the cellulose content of the moxa floss with unknown leaf-fuzz ratio are in, the leaf-fuzz ratio of the moxa floss can be obtained, that is, the quality of the moxa floss with unknown leaf-fuzz ratio is identified.

The invention discloses pseudomonas fluorescens SMP5 which is derived from rhizosphere soil of salvia miltiorrhiza. The invention further provides application of the pseudomonas fluorescens SMP5 in growth promotion and quality improvement of the artemisia annua, the pseudomonas fluorescens SMP5 obtained through screening and identification can achieve efficient colonization on the surfaces of leaves of the artemisia annua, and plant growth and quality improvement are remarkably promoted in a foliage spraying mode. Compared with an existing traditional method depending on chemical fertilizer or exogenous hormone, the strain can effectively promote leaf yield and biomass accumulation (the yield and the biomass are increased by 41.04% and 44.92% respectively), the secretory glandular hair density is remarkably increased by 4.4 times, and more structural bases are provided for artemisinin synthesis. According to the method, a complete regulation and control chain of beneficial bacterium-leaf surface-glandular hair-artemisinin synthesis is established, double improvement of quality and yield is achieved, and the method has the advantages of being environmentally friendly, safe and capable of being popularized.

The invention discloses an application of plant epidermal hair instability in increasing the iron content of plant leaves. Belongs to the technical field of plant nutrient enrichment. The invention provides a method for increasing the iron content in a plant leaf, and the iron content in the plant is increased by destabilizing epidermal hair. The method is suitable for plants with non-glandular hair distributed on leaves such as arabidopsis thaliana and Chinese cabbage, hunger response of iron ions in the leaves is caused through methods such as leaf epidermal hair touch treatment, then the iron content in the plant leaves is increased, and the method has the advantages of being short in time, simple and easy to operate.

The application relates to the technical field of plant molecular biology, and particularly discloses a tomato gland-type epidermal hair regulating gene SlWRKY42 and application thereof, wherein the ORF sequence of the gene SlWRKY42 is shown as SEQ ID NO. 1. By regulating the overexpression of the SlWRKY42 gene in a tomato plant, the density of tomato gland hairs is significantly increased, the transcription and expression of an eneyterpene synthase gene (SlTPS12) and a precursor synthesis pathway transgene are significantly up-regulated, and the content of eneyterpene substances is significantly increased. The gene SlWRKY42 provided by the application can regulate the growth and development of tomato gland-type epidermal hairs, and simultaneously regulate the synthesis of eneyterpene substances.

The present invention relates to a method of producing a plant with an increased density of glandular trichomes and / or with an increased production of metabolite(s) comprising a step of introducing at least one genetic alteration that increases activity of PRODERMAL FACTOR2 (PDF2) of said plant, wherein PDF2 preferably has an amino acid sequence selected from the group consisting of SEQ ID NO: 1, SEQ ID NO: 2, SEQ ID NO: 3, SEQ ID NO: 4, SEQ ID NO: 5, SEQ ID NO: 6, SEQ ID NO: 7, SEQ ID NO: 8, SEQ ID NO: 9, SEQ ID NO: 10, SEQ ID NO: 11, SEQ ID NO: 12, SEQ ID NO: 13, SEQ ID NO: 14, SEQ ID NO: 15, SEQ ID NO: 38, SEQ ID NO: 39, SEQ ID NO: 60, SEQ ID NO: 62, SEQ ID NO: 64, SEQ ID NO: 66, SEQ ID NO: 68, SEQ ID NO: 70, SEQ ID NO: 72, SEQ ID NO: 74, SEQ ID NO: 75, SEQ ID NO: 77, SEQ ID NO: 79, SEQ ID NO: 81, SEQ ID NO: 83, SEQ ID NO: 85, SEQ ID NO: 87, SEQ ID NO: 89 and SEQ ID NO: 90.

The invention discloses a mint McHD-Zip1 gene promoter and application thereof, and relates to the technical field of gene engineering. According to the mint McHD-Zip1 gene promoter disclosed by the invention, the nucleotide sequence of the mint McHD-Zip1 gene promoter is as shown in SEQ ID NO. 1. When the promoter is used for constructing a plant expression vector pGC-GUS-McHD-Zip1pro, the result of the embodiment shows that the promoter can drive a target gene to be predominantly expressed in stem tips, leaf axils and secretory glandular hairs of a plant instead of being widely expressed in various tissues and organs of the whole plant, so that the adverse effect on normal growth and development of the plant can be reduced, and the plant expression vector pGC-GUS-McHD-Zip1pro can be used for preparing the plant expression vector pGC-GUS-McHD-Zip1pro. The metabolism or development process of the specific part of the plant can be more accurately regulated and controlled; compared with a constitutive promoter, the specific promoter avoids interference of whole plant expression on plant metabolic balance, and provides a precise tool for studying plant glandular hair development and metabolic engineering.

An electrostatic separation apparatus and method are provided for isolating glandular trichomes from trichome-bearing plant biomass such as cannabis, hemp, or hops. The system may include a vertically oriented particle transport assembly configured to convey a sample through a linear free-fall pathway, a separation chamber with opposing electrode assemblies generating a uniform electrostatic field, and discharge electrodes positioned below the chamber to neutralize residual charge. Airflow regulation may be achieved through laminar diffuser plates and periodic vibration of critical components, including the hopper and diffuser, to prevent clogging and maintain uniform flow. The hopper may further include a circular discharge spout and rounded internal fillets to prevent bridging and ensure consistent powder recirculation. The apparatus achieves high-purity trichome separation through controlled field exposure, aerodynamic stabilization, and charge-neutralized particle discharge, enabling continuous, high-throughput operation with improved yield, reduced maintenance, and enhanced reproducibility across successive processing cycles.

The present invention discloses a plant glandular hair head gene GHF1 The present invention identifies a gene that can negatively regulate the development of plant glandular hair heads. GHF1 , constructed its overexpression vector and gene editing vector, and carried out tobacco transformation respectively to obtain the corresponding overexpression strains and gene knockout plants; the epidermal hair morphology identification and leaf chemical composition analysis showed that GHF1 The gene can negatively regulate the occurrence and development of the glandular hair head of plants, and has important application value in the regulation of glandular hair development and the synthesis of glandular hair-specific substances.

A method for identifying the authenticity of moxa based on the fluorescence microscopic characteristics of non-glandular hairs, comprising the following steps: (1) weighing the sample; (2) treating with formaldehyde solution; (3) rinsing; (4) treating with 2-phenylboronic acid 2-aminoethyl ester reagent; (5) microscopic observation: dripping glycerol on the moxa, observing under ultraviolet light excitation of a fluorescence microscope, judging the authenticity of the moxa according to the microscopic characteristics, if the T-shaped non-glandular hairs are observed to have handles or shedding marks of varying lengths, it is true moxa, otherwise it is false moxa. The method of the present invention is simple and easy to operate. Under the action of formaldehyde and 2-phenylboronic acid 2-aminoethyl ester, the top cells of the real moxa after mechanical processing regain fluorescence, and the stem cells are also strengthened. It can be clearly seen that the T-shaped non-glandular hairs in the moxa have handles or shedding marks of varying lengths, while the fake moxa does not have the handles or shedding marks unique to the T-shaped non-glandular hairs. The authenticity of the moxa is effectively solved, the treatment and efficacy are ensured, and the economic and social benefits are huge.

The invention discloses an application of a cucumber CsLSGT gene as a target spot in regulating formation of a silicification layer on the surface of a fruit gland hair. The formation of the silicification layer on the surface of the cucumber gland hair is reduced by reducing the expression quantity of the CsLSGT gene. It is found through screening that the cucumber CsLSGT gene is related to formation of a silicification layer on the cucumber fruit gland hair surface. In a cucumber mutant, the silicon element content of fruit glandular hair is obviously reduced compared with that of a wild type '649'. Meanwhile, a grafting experiment shows that the function of the CsLSGT gene depends on the overground part, which indicates that the reduction of the silicon element on the surface of the glandular hair is mediated by an overground part factor. The gene can be used for screening cucumber fruits with few silicon elements and improving the commodity quality traits of the cucumber fruits.

The invention discloses a method for assisted breeding of a high-southern-rust-resistance corn inbred line by using leaf glandular hair traits, and relates to the technical field of crop genetic breeding, and the method comprises the following specific steps: hybridizing the high-southern-rust-resistance corn inbred line with a to-be-improved southern-rust-susceptible corn inbred line or hybridizing and then backcrossing to select the line, selfing F1 generation and reserving seeds, and breeding the F1 generation and the F1 generation to obtain the high-southern-rust-resistance corn inbred line. Starting from the F2 generation or the BC1F1 generation, selecting plants or rows with large leaves and fewer hairs according to the form and the quantity of glandular hairs of the leaves, selecting single plants with good corn characters, and continuously selfing or backcrossing for N generations, so as to obtain the high-southern-rust-resistance corn selfing line with stable inheritance. According to the method disclosed by the invention, the corn inbred line with high southern rust resistance is bred according to the quantity and the sparse degree of the big hairs in the corn leaf glandular hairs, so that multiple limitations of inoculation identification and special habitat conditions can be overcome, the breeding process of the corn inbred line with high southern rust resistance is accelerated, accurate identification of resistant materials is realized, and a foundation is laid for cultivation of new varieties.

The invention relates to a preparation method of a scanning electron microscope sample for observing microscopic characteristics of the lower surface of a folium artemisiae argyi plant, and can effectively solve the problems that the epidermis is difficult to tear and dissociate due to the fact that the folium artemisiae argyi plant is a paper leaf and is thin, even if non-glandular hair on the lower surface can be successfully removed and the epidermis can be separated out, the epidermis cannot be separated out, and the microscopic characteristics of the lower surface of the folium artemisiae argyi plant cannot be observed. In order to solve the problem that the microscopic characteristics of the stem and the glandular hair of the T-shaped non-glandular hair are difficult to observe due to the removal of the glandular hair and the whole T-shaped non-glandular hair, the technical scheme for solving the problem is as follows: the method comprises the following steps of: 1) shearing leaves; 2) methanol fixation; the preparation method comprises the following steps of 1, sample preparation, 2, sample preparation, 3, chloral hydrate treatment, 4, phosphate buffer solution rinsing, 5, pectinase solution treatment, 6, ultrasonic treatment, 7, rinsing and 8, sheet sealing. The preparation method is simple in operation, low in cost, small in workload, high in success rate and clear in observation, observation of microscopic characteristics of the lower surfaces of various folium artemisiae argyi plants can be successfully achieved, and the preparation method is an innovation of the preparation method of the artemisia folium artemisiae argyi plant structure observation sample.

This invention discloses a gene regulating plant glandular trichome development. NtJAZ11, This application aims to address the current technical problem of unclear developmental mechanisms and lack of effective regulatory methods and utilization pathways for glandular trichomes and protective trichomes in plants (such as tobacco). The gene encoding this gene, its expression vector, and its applications are described. NtJAZ11 Overexpression and gene-editing vectors were constructed, and corresponding overexpression and gene-editing lines were obtained through transformation. Subcellular localization, tissue expression, MeJA response pattern analysis, and plant phenotypic identification showed that... NtJAZ11‑2 It can negatively regulate the development of tobacco glandular hairs, and has important application value in regulating the development of plant epidermal hairs.

This invention application discloses a gene regulating plant glandular trichomes. NtMIXTA This application relates to genes, their encoded proteins, biological materials, and applications, aiming to address the technical problem of low efficiency in increasing plant glandular trichome density using traditional breeding techniques. This application clones a gene from tobacco... NtMIXTA Genes, and constructed separately NtMIXTA Gene knockout and overexpression lines were found NtMIXTA Gene knockout reduced leaf glandular hair density, while overexpression lines showed a significant increase in leaf glandular hair density, indicating that... NtMIXTA Genes positively regulate the occurrence and development of glandular trichomes in tobacco. This application is of great significance for studying the molecular regulatory network of plant glandular trichome formation.

The invention discloses application of an NtHCH5 gene in regulation and control of plant glandular hair development, and belongs to the technical field of plant genetic engineering. The nucleotide sequence of the gene for regulating and controlling plant glandular hair development provided by the invention is as shown in SEQ ID NO. 1. An RNAi carrier of the NtHCH5 gene is constructed through an RNAi technology, a tobacco plant silenced by the NtHCH5 gene is cultivated, and it is found that compared with a normal tobacco plant, the density of glandular hairs in tobacco leaves after the NtHCH5 gene is silenced is remarkably reduced, and the glandular hairs are shortened. An economic plant tobacco is used as a research object, and experiments prove that the NtHCH5 gene participates in regulation and control of plant glandular hair development, a new target gene is provided for regulation and control of plant glandular hair development, and a gene regulation and control network of plant in-vivo epidermal hair is enriched. The gene function of the plant NtHCH5 gene is also enriched, and the gene has important significance for analyzing the diversity of the plant NtHCH5 gene.

The invention discloses a plant glandular hair head generation gene GHF1 as well as an expression product, an expression vector and application thereof, and aims to solve the technical problems of insufficient research on the glandular hair head generation gene and unclear glandular hair generation molecular regulation network at present. According to the invention, a gene GHF1 capable of negatively regulating plant glandular hair head generation is identified, an overexpression vector and a gene editing vector of the gene GHF1 are constructed, and tobacco transformation is respectively carried out to obtain a corresponding overexpression strain and a gene knockout plant; epidermal hair morphological identification and foliar chemical component analysis show that the GHF1 gene can negatively regulate and control plant glandular hair head generation and development, and has important application value in glandular hair development regulation and control, glandular hair specific substance synthesis and the like.

The invention particularly discloses a method for inducing transdifferentiation of paving cells in mature leaves of nicotiana benthamiana into glandular hair cells, which comprises the following steps: preparing an agrobacterium infection solution, and agrobacterium is selected from one or more of a GV3101 strain, a C58C1 strain or a K599 strain; carrying out infection treatment on completely expanded mature leaves of the nicotiana benthamiana by using an agrobacterium infection solution; and after infection, the nicotiana benthamiana is cultured, and the paving cells of the leaves are induced to be transdifferentiated into glandular hair cells. The method is easy to operate and good in repeatability, stable genetic transformation and strict sterile conditions are not needed, the paving cells of the Bensi tobacco leaves can be induced to be transdifferentiated into the glandular hair cells, and increase of the number of the glandular hair cells is promoted.