CRISPR-SaCas9 gene editing system and application thereof
A technology for gene editing and encoding genes, applied in gene therapy, genetic engineering, DNA/RNA fragments, etc. It can solve the problem of low gene editing efficiency, and achieve the effect of improving gene knock-in efficiency, low off-target efficiency and broad application prospects.
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Embodiment 1
[0041] Embodiment 1 vector construction
[0042] This example is used to construct each vector of the gene editing system described in the present invention.
[0043] (1) Construction of different nuclear localization signal SaCas9 vectors
[0044] Different nuclear localization signal sequences were obtained by PCR amplification, and then EF1 promoter, different nuclear localization signal sequences (SEQ ID NO.8-14), SaCas9 (encoded by SEQ ID NO. The nucleotide sequence of .15), Wpre and PolyA were spliced and cloned into the pEF1-SaCas9 plasmid; wherein, SaCas9 and the nuclear localization signal were connected by connecting sequence 1 (SEQ ID NO.16); digested with endonuclease and Sanger sequencing identified SaCas9 vectors with different nuclear localization signals.
[0045] (2) Construction of sgRNA vectors with different backbone structures and different target sequence lengths
[0046] Use the CHOPCHOP website (https: / / chopchop.rc.fas.harvard.edu / ) to design sgRNA...
Embodiment 2
[0051] Example 2 Gene editing method of human induced pluripotent stem cells at different sites
[0052] This example provides gene editing methods of human induced pluripotent stem cells at different sites.
[0053] (1) Electroporation of human induced pluripotent stem cells
[0054] Human induced pluripotent stem cells using mTeSR TM 1 culture medium. During recovery and passage, the culture plate was first coated with Matrigel. The specific steps are as follows: Before electroporation, iPSC cells were digested with Accutase to obtain a single cell suspension; 0.8-1.5×10 6 The cells were electroporated, and the amount of gene editing plasmids was as follows: 1 μg of Cas9, 0.5 μg of sgRNA (cutting the genome), 0.5 μg of sgDocut (cutting of pDonor), 1 μg of pDonor and 0.5 μg of BCL-XL plasmid; Human Stem Cell was used Kit 2 (Lonza) was used for electroporation, and the electroporation procedure was B-016.
[0055] (2) Gene editing of human induced pluripotent stem cells ...
Embodiment 3
[0063] Example 3 SaCas9 carrying nuclear localization signal improves gene editing efficiency
[0064] This example is used to compare the effects of different nuclear localization signals on the efficiency of SaCas9 gene editing. SaCas9 plasmids with different nuclear localization signals, including: Group 1: SaCas9 (No NLS) (1 μg), Group 2: SaCas9-NPM-NLS (1 μg), Group 3: SaCas9-BPNLS (1 μg), Group 4: BPNLS-SaCas9-BPNLS (1 μg) and Group 5: HMGA2-SaCas9-BPNLS (1 μg) (eg image 3 shown), co-electroporated iPSCs with CRISPR components such as pU6-sgEEF1A1 (0.5 μg), pD-EEF1A1-E2A-mNeonGreen-sg (1 μg), pU6-sgDocut (0.5 μg) and pEF1-BCL-XL (0.5 μg) , The gene knock-in efficiency was analyzed by FACS on the 3rd day after electroporation.
[0065] Such as Figure 4 and Figure 5 As shown, the HDR efficiency at EEF1A1 increased from 17% of the control SaCas9 (No NLS) to 30.2% (SaCas9-NPM-NLS), 29.9% (SaCas9-BPNLS), 25.8% (BPNLS-SaCas9-BPNLS) and 41 %(HMGA2-SaCas9-BPNLS). These ...
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