System for rapid production of high-titer and replication-competent adenovirus-free recombinant adenovirus vectors
A recombinant adenovirus, adenovirus technology, applied in the fields of immunology, gene therapy and vaccine technology
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Embodiment 1
[0134] Example 1: Generation of Replication Defective Adenovirus Expressing Influenza HA
[0135] Two adenoviral (Ad) vectors encoding influenza HA were constructed using the AdEasy system. The 2 most recent influenza strains [A / Panama / 2007 / 99 (H3N2) and B / Hong Kong / 330 / 01] selected for vaccine production in 2003-2004 were reviewed by The Centers for Disease Control and Prevention. Prevention) (CDC). A / Panama / 2007 / 99 HA was cloned by reverse transcription of influenza RNA followed by amplification of the HA gene by polymerase chain reaction (PCR) using the following primers:
[0136] Table 1: Primer sequences used to amplify influenza genes
[0137]
[0138] These primers contained sequences that anneal to the 5' and 3' ends of the A / Panama / 2007 / 99 HA gene, the eukaryotic ribosome binding site immediately upstream of the HA start ATG codon (Kozak, 1986), and the KpnI site for subsequent cloning. The KpnI fragment containing the full-length HA gene was inserted in the ...
Embodiment 2
[0142] Example 2: Construction of pAdHighα
[0143] Crucell's shuttle plasmid pAdApt was digested separately with restriction enzymes SgrAI+EcoRI, and BstXI+EcoRI. In parallel, the shuttle plasmid pShuttleCMV was digested with SgrAI+BstXI. The resulting pAdApt SgrAI-EcoRI and BstXI-EcoRI fragments were inserted into the SgrAI-BstXI site of pShuttleCMV by 3-route ligation, resulting in a replication-deficient Ad vector. A replication-defective Ad vector encoding the influenza HA gene (Ad Highα PNM2007 / 99.H3) was produced by transfecting the recombinant plasmid into PER.C6 cells. The cytopathic effect (CPE) appeared around 7 days after infection, the same time frame as that required for the AdEasy system in 293 cells (He et al., 1998).
Embodiment 3
[0144] Embodiment 3: Construction of pAdHighβ
[0145] To repair the defective sequence, the CMV promoter, adjacent multiple cloning site, and flanking Ad sequences of pShuttleCMV as 1 unit were replaced by their counterparts from pAdApt by homologous recombination, since the 2 shuttle plasmids share extensively overlapping sequences. However, new markers are also required for selection of recombinants. The full-length tetracycline (Tc) resistance gene from plasmid pBR322 (Backman and Boyer, 1983; Peden, 1983) was performed by PCR using primers 5′-GAGCTCGGTACCTTCTCATGTTTGACAGCTTATCAT-3′ and 5′-TCTAGAGGTACCAACGCTGCCCGAGATGCGCCGCGT-3′ with a built-in KpnI site Amplify. The amplified Tc gene was inserted into the KpnI site of the Amp resistance plasmid pAdApt to generate a new plasmid pAdApt-Tc, which can be selected by application of Amp and Tc to the growth medium.
[0146] The Ad sequence in pShuttleCMV was replaced with its counterpart in pAdApt-Tc using the high efficien...
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