Mait-like cells and method for manufacturing same
A production method and cell therapy technology, applied in animal cells, vertebrate cells, genetically modified cells, etc., can solve problems such as difficult animal function analysis, unknown cell lines with this characteristic, and lack of cell proliferation ability
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Embodiment 1
[0104] Example 1: Establishment of iPS cells from human-derived MAIT cells
[0105] Mononuclear cells were prepared from human cord blood using Ficoll. The mononuclear cells were mixed with a biotin-labeled substance of the monoclonal antibody 3C10 (gifted by Dr. Olivier Lantz of L'Institut Curie, France, or Biolegend) that specifically recognizes TCR (Vα7.2) of MAIT cells, Cells reactive with the 3C10 antibody were positively selected using a MACS column (manufactured by Miltenyi Biotech) using avidin magnetic beads to concentrate MAIT cells. This operation was performed using umbilical cord blood from 3 different donors. As a result, the purity of MAIT cells defined as 3C10 positive cells by FACS analysis was 96%, 88%, and 78%, respectively.
[0106] The 200,000 3C10-positive cells thus purified were infected with SeVdp(KOSM)302L (gifted by Dr. Makoto Nakanishi from the Institute of Advanced Industrial Research) serving as a vector for production of human iPS cells for 2 ...
Embodiment 2
[0111] Example 2: Characteristic analysis of MAIT-iPS cells
[0112] The MAIT-iPS cell lines 1-3D, 2-5D, and 4-6D individually established and isolated in Example 1 showed a single layer, flat outline, and clear colonies on MEF feeding, which showed that they were different from ordinary human ES / iPS cells. Cells with very similar morphology ( Figure 2A ).
[0113] In addition, the expression of markers specific to human ES / iPS cells was investigated using MAIT-iPS cells thus subcultured. For fixed MAIT-iPS cells, anti-alkaline phosphatase (ALP) antibody, anti-SSEA4 antibody, anti-Oct-3 / 4 antibody, anti-Nanog antibody (above, R&D Systems Inc.), anti-TRA-1 antibody as primary antibody -60 antibody (BD Bioscience) or anti-TRA-1-81 antibody (Santa Cruz) was reacted, and rhodamine-labeled secondary antibody (Jackson ImmunoResearch) was used for staining. Nuclei were stained with 4',6-diamidino-2-phenylindole (DAPI: 4',6-diamidino-2-phenylindole) solution (1 µg / mL). The stai...
Embodiment 3
[0133] Example 3: Production of MAIT cells from MAIT-iPS cells
[0134] Regarding stem cells such as ES cells, it is known that by co-culturing OP9 cells as feeder cells, differentiation can be induced into CD34-positive blood cell / lymphocyte progenitor cells, and further, by co-culture with OP9 cells that forcibly express Notch ligand DLL1 (OP9 / DLL1) co-culture, can differentiate into cells of T cell series (Schmitt TM et al., Nat. Immun.5, 410-417 (2004); Wakao H et al., FASEB J.22, 2223-2231 ( 2008); Wakao H et al., WO2008 / 038579; Watarai H et al., J.Clin.Invest.120, 2610-2618 (2010); WO2010 / 027094; Timmermans F et al., J.Immunol.182, 6879-6888 (2011)).
[0135] In this example, induction of differentiation from MAIT-iPS cells into MAIT cells was attempted based on this method.
[0136] As OP9 cells and OP9 / DLL1 cells, cells purchased from Riken Cell Bank were used. First, on the OP9 cells 3 to 7 days after fusion, a substance that disperses the colony of MAIT-iPS cell...
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