A gene base editor
By fusing human cytosine deaminase 3A with the CRISPR/Cas system and combining it with a uracil glycosylase inhibitor, efficient base editing at the GpC site is achieved, solving the problem of HDR-mediated gene correction in the existing technology. The problems of low efficiency and inability to edit GpC sites have improved the accuracy and application scope of gene editing.
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[0084] Example 1: Base editor
[0085] The expression plasmid of pCMV-hA3A-BE was constructed. Human apolipoprotein B messenger RNA deaminase catalytic subunit 3A (APOBEC3A, hA3A; SEQ ID NO: 1) with Cas9 nickase and a uracil DNA glycosidase inhibitor [Bacillus phage] (SEQ ID NO: 12 ) fused to an expression vector. The 10th aspartic acid of Cas9 nickase is mutated to alanine, thereby losing the activity of cutting double strands and ensuring a nick on one strand.
[0086] The fusion expression vector hA3A-nCas9-UGI (hA3A-BE, SEQ ID NO: 21) and the expression vector of single-stranded guide RNA were co-transformed into eukaryotic cells ( figure 1 , legend A), C-T base editing occurs at the site targeted by the guide RNA of the genome. The sequence of the genomic DNA target position was amplified by PCR, and the base editing efficiency of the target site C-T was detected by Sanger DNA sequencing. Compared with the co-expression of sgRNA and BE3, the method of co-expression of...
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