An Orthogonal Aminoacyl-tRNA Synthetase/tRNA System Using a Chimeric Design Approach
A design method, a technology for synthesizing enzymes, applied in the field of chemical biology, which can solve the problems of increasing restrictions, not widely orthogonal, and low activity of unnatural amino acids
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Embodiment 1
[0065] Embodiment 1: Construction of chimeric histidine tRNA
[0066] In the present invention, the chimeric tRNA and the reporter gene GFP190TAG-His6 are constructed on the same plasmid, controlled by the lpp promoter and the pBAD promoter respectively. The specific construction method is as follows:
[0067] (1) Construct GFP190TAG-His6 onto the pNEG vector
[0068] The sequence of GFP190TAG-His6 is shown in SEQ ID No.1. Primers pNEG-gfp-F and pNEG-gfp-R were designed to amplify using the existing plasmid containing this gene as a template, and pNEG-gfp-v-F and pNEG-gfp-v-F were simultaneously designed and synthesized. pNEG-gfp-v-R, using the previous pNEG as a template to amplify the vector. After the agarose gel was recovered, the Gibson assembly was used to transform DH10B competent cells, and the single clone was selected and sequenced to obtain the plasmid pNEG-GFP190TAG-His6;
[0069](2) Design chimeric histidine tRNA and clone into pNEG-GFP190TAG-His6 vector
[00...
Embodiment 2
[0073] Embodiment 2: Construction of chimeric histidyl-tRNA synthetase
[0074] The chimeric histidyl-tRNA synthetase is composed of two parts, one is the pyrrolysyl-tRNA synthetase tRNA binding domain, and the other is the catalytic domain of the histidyl-tRNA synthetase, see figure 2 and 3 c.
[0075] The construction of the chimeric histidyl-tRNA synthetase includes the selection of the tRNA binding domain of the pyrrolysyl-tRNA synthetase, the selection of the catalytic domain of the histidyl-tRNA synthetase, and the selection of the fusion mode of the two.
[0076] (1) Analysis and selection of the tRNA binding domain of pyrrolysyl-tRNA synthetase
[0077] In the present invention, it is necessary to first clone the pyrrolysyl-tRNA synthetase tRNA binding domain onto the pBK carrier. This tRNA binding domain can be divided into two subdomains, and the N-terminal 1-149 recognizes the variable region of the corresponding tRNA. And the T loop, the 185-240 section recogni...
Embodiment 3
[0084] Example 3: Optimization of chimeric histidyl-tRNA synthetase linker peptide
[0085] In the construction of chimeric protein, the connecting peptide between two chimeric fragments is crucial to the activity of chimeric molecule, in the present invention we have tested the length of connecting peptide and the kind of connecting peptide ( Figure 11 ).
[0086] (1) Optimization of connecting peptide length
[0087] The length of the designed flexible connecting peptide (GS-type) is 0, 6, 12, 18 and 24 amino acids, named chHisRS4-2, -3, -4, -5 and -6, respectively, and the nucleotide sequence of the corresponding connecting peptide See Table 3-1. At the same time, the N-terminal 15 amino acids of histidine-tRNA synthetase were deleted on the basis of the 0 amino acid linking peptide, and named chHisRS4-1. The designed primers were constructed on the vector by the Q5 site-directed mutagenesis kit (NEB), corresponding The primers are listed in Table 3-2.
[0088] (2) Con...
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