Mitochondrion RNA polymerase subsidiary transcription factor as well as polymerase-transcription factor compound and application thereof
An RNA polymerase and transcription factor technology, applied in the field of novel polymerase-transcription factor complexes, which can solve problems such as mitochondrial DNA deletion
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Embodiment 1
[0076] Example 1: Cloning of novel mitochondrial RNA polymerase Rpo41 and accessory transcription factor Mtf1 from fission yeast
[0077] According to the amino acid sequence of the homologous protein in budding yeast, Blast comparison was carried out on Genedb, the website of the Sanger Research Center in the United Kingdom. The results showed that there was a highly conserved homologous mitochondrial RNA polymerase in fission yeast, and the system name was SPAC26H5.12 , and a highly conserved mitochondrial transcription factor, the system name SPAC1002.08c. The amino acid sequence comparison results between Mtf1 homologous proteins in fission yeast and budding yeast can be found in figure 1 , by comparison, the applicant found that fission yeast Mtf1 has a conserved N-terminal domain and a conserved C-terminal domain, see figure 2 .
[0078] In the following embodiments of the present invention, unless otherwise specified, SPAC26H5.12 is referred to as Rpo41 for short, an...
Embodiment 2
[0079] Example 2: Sublocalization of fission yeast Mtf1 in cells
[0080] In order to study the function of fission yeast Mtf1 in cells, the present invention uses the ultracentrifugation method to extract fission yeast mitochondria, nucleus and cytoplasmic components respectively, and uses the Western-blot method to detect the sublocalization of Mtf1 in cells, such as Figure 4 As shown, Mtf1 is localized in both the nucleus and mitochondria. This result suggests that Mtf1 not only functions in the mitochondria of fission yeast, but also has the function of transcribing genes in the nucleus.
Embodiment 3
[0081] Example 3: Fission yeast Mtf1 knockout mutants have defects in cell cycle regulation and cell morphology
[0082] In order to explore its cell function, the present invention knocks out the Mtf1 gene of fission yeast. The knockout is carried out in diploid cells, and the spores produced by mating are observed under a microscope after germination. Cells lacking Mtf1 exhibited a similar morphology, ie, the knockout strain failed to form colonies and could only divide a few times on the plate. image 3 showed that cells lacking Mtf1 were morphologically longer and larger than wild-type cells, and became egg- or bottle-shaped. DAPI staining showed that Mtf1-deficient cells had no apparent nuclear defects. Staining with fluorescent white Calcofluor (Calcofluor fluorescent stain) showed that cells lacking Mtf1 had a higher proportion of cell compartments, some of which contained multiple cell compartments. This result suggests that Mtf1 may have the function of regulating c...
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