A base editing system based on Streptococcus pyogenes and its application in gene editing
A Streptococcus pyogenes, base editing technology, applied in the field of molecular biology, can solve problems such as hindering the application of base editing technology
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Embodiment 1
[0048] Example 1 Preparation method of base editing system components based on Streptococcus pyogenes
[0049] Figures 1 to 4 Expression vector maps for more precise base editing systems for four Streptococcus pyogenes.
[0050] This example provides the preparation method of rAPOBEC1:Cas9:UGI mRNA and gRNA based on the base editing system components of Streptococcus pyogenes.
[0051] 1. Preparation of Streptococcus pyogenes rAPOBEC1:Cas9:UGI mRNA, the method is as follows:
[0052] (1) Preparation of transcription vector for HF1-BE2, HF1-BE3, HF2-BE2 or HF2-BE3, the sequence is shown in SEQ ID NO.3, SEQ ID NO.4, SEQ ID NO.5, SEQ ID NO.6 .
[0053] (2) Prepare a transcription template of Streptococcus pyogenes APOBEC1:Cas9:UGI containing the T7 promoter:
[0054]The transcription vector of HF1-BE2, HF1-BE3, HF2-BE2 or HF2-BE3 was cleaved with KpnI, and then the cleaved product was subjected to column purification with a PCR product purification kit (Axygen), followed by ...
Embodiment 2
[0066] Example 2 Preparation of rAPOBEC1:Cas9:UGI mRNA and gRNA components of the base editing system based on Streptococcus pyogenes
[0067] Specifically, the operation procedure of the preparation method of Streptococcus pyogenes APOBEC1:Cas9:UGI mRNA, APOBEC1:Cas9:UGI protein and gRNA described in Example 1 is as follows:
[0068] 1. Preparation of APOBEC1:Cas9:UGI mRNA and gRNA transcription template DNA:
[0069] (1) Preparation of S. pyogenes APOBEC1:Cas9:UGI mRNA transcription template DNA
[0070] The transcription vector of HF1-BE2, HF1-BE3, HF2-BE2 or HF2-BE3 was prepared by plasmid extraction (independently developed), and then the transcription vector was digested with KpnI, and the reaction system was carried out according to the following table 1:
[0071] Table 1 Reaction system
[0072] Element Dosage mRNA transcription vector 2000ng 10X NEBuffer 1.1 5μl Kpn I 5μl ddH 2 O
Make up to 50μl
[0073] Digestion ove...
Embodiment 3
[0133] Example 3 Site-directed mutagenesis of Tyr gene mediated by a more precise base editing system based on Streptococcus pyogenes
[0134] 1. In order to use a more precise base editing system based on Streptococcus pyogenes to achieve single-gene site-directed mutagenesis in mouse fertilized eggs, we designed two gRNAs (gRNA-1 and gRNA-2) for the Tyr gene. Site-directed mutagenesis of the Tyr gene results in the formation of a stop codon, which results in premature termination of the Tyr gene translation, which changes the coat color of the pups from black to white.
[0135] First, we transcribed the Tyr gRNA, and then mixed Tyr gRNA (50ng / μl) with rAPOBEC1:Cas9:UGI(HF2-BE2) mRNA (100ng / μl) and injected it into 0.5-day-old mouse fertilized eggs. The site-directed mutation efficiency was detected 48 hours after injection. By combining PCR and Sanger sequencing, we found that for gRNA-1, 11.6% of mouse embryos were mutated, while for gRNA-2, 50% of embryos were mutated. . ...
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