Expression and preparation method of granulysin
A technology of granullysin and expression cassette, which is applied in the field of expression and preparation of granullysin, and can solve the problems of affecting biological activity, difficulty in renaturation, and low expression efficiency
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Embodiment 1
[0181] Embodiment 1, the construction of GNLY expression vector
[0182] The full length of GNLY is 145 amino acids, including the predicted signal peptide, which is located at amino acid positions 1-22. In killer T cells and NK cells, GNLY is synthesized in the form of 15kDa and then cleaved at the first position to form the form of 9kDa (such as figure 1 A).
[0183] The inventors connected the coding sequence of 9kDa and 15kDa GNLY to the 3' end of the α-Factor signal peptide of Saccharomyces cerevisiae, and only kept the restriction site of kex2 protease, and finally cloned it into pPIC9K yeast through BamH I and EcoR I restriction sites in the expression vector. In this way, methanol induces the expression of AOXI promoter to drive the expression of its downstream proteins. Schematic diagram of the structure of GNLY expression vector figure 1 Shown in B.
Embodiment 2
[0184] The screening of embodiment 2, GNLY expression bacterial strain and shake flask expression condition optimization
[0185] After linearizing pPIC9K-GNLYs-His with SalI, it was electroporated into Pichia pastoris GS115 strain. Nine clones were selected from the YPDG plates where the 9kDa and 15kDa GNLY expression strains were cultivated for induced expression. Western blotting was used to detect the expression of GNLY in the culture supernatant of these clones.
[0186] Taking the expression of 9kDa GNLY clones as an example, most of the clones effectively express 9-kD GNLY (such as figure 2 A), the clone with the highest expression was named GS115 / 9kDa GNLY-His. Western blotting showed that the molecular weight of 9kDa GNLY expressed by yeast was between 10-15kDa, and there were two bands (15% gel) with very similar molecular weights (different degrees of glycosylation modification). The expression level of 15kDa GNLY was significantly higher than that of 9kDa GNLY,...
Embodiment 3
[0188] Embodiment 3, the fermentation expression of GNLY pilot scale
[0189] The present inventors cultivated the expression strain GS115 / 9-kD GNLY-His by fed-batch cultivation (fed-batch cultivation) method, and optimized the method flow. The inventors first used a basal salt medium (pH 5.0) containing 4% (v / v) glycerol to amplify the bacteria in the batch stage (the pH was subsequently adjusted to maintain pH 5.0), and the culture temperature was 30°C. When the DO rises sharply (about 22 hours of cultivation), the batch stage ends, and the wet weight of the bacteria reaches 118.7g / L. Then it enters the conversion period, that is, the glycerin feeding period (feeding amount is: 18.15ml / hr / liter, in the feeding glycerin, each liter of 50% w / v glycerin solution contains 12ml of PTM1, PTM1 is a kind of yeast growth needs Trace element salt solution, the composition is: copper sulfate-5H 2 O 6.0g, sodium iodide 0.08g, manganese sulfate-H 2 O 3.0g, sodium molybdate-2H 2 O 0.2...
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