DNA system for selectively killing cells by inducing multi-site cutting genome by utilizing RNA interference mechanism
An RNA interference and genomic technology, applied in the field of DNA systems, can solve the problems of complex redundancy and correlation, side effects, and no obvious regulatory effect.
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Embodiment 1
[0070] Materials and methods
[0071] Construction of plasmids and vectors
[0072] Cas9 was ligated into a piggyBac vector containing a hygromycin resistance gene driven by a PGK promoter driven by a CAGGS promoter. The pre-sgRNA (sgRNA sequence containing RNA sequences completely complementary to the target miRNA or siRNA at both ends, referred to as miRT-sgRNA-miRT) was connected to the bleomycin (zeocin) resistance gene driven by the PGK promoter. In the piggyBac vector, the pre-sgRNA is also driven by the CAGGS promoter. In order to prepare the miRT-sgRNA-miRT construct sensitive to target miRNA or siRNA, use Fastpfu polymerase (Transgene Company), with the plasmid containing sgRNA as a template (the sequence of the sgRNA is complementary to the partial sequence of the Alu element), with Primers corresponding to miRT were used for PCR. The PCR product was incubated with EcoRI-HF (20U, NEB Company) and BamHI-HF (20U, NEB Company) at 37°C for 2h, purified with a spin c...
Embodiment 2
[0083] To test the killing effect of MICR-CUT system on Hela cells.
[0084] In this study, the Alu element with a high repetitive sequence in the Hela cell genome was first selected, and two sgRNAs targeting the Alu element were designed. -OFFinder software forecast). In this study, these two sgRNAs were constructed as pre-sgRNAs (referred to as 294T-Alu1, 294T-Alu2) that can be activated by miR-294. See the experimental results figure 2 , indicating that Cas9 with nuclease activity and pre-sgRNA expression vectors were transfected into Hela cells, and both pre-sgRNAs could cause about 50% reduction in the number of Hela cells under the condition of miR-294 transfection. The experimental results preliminarily indicate that the MICR-CUT DNA system can be used to specifically kill cells by targeting and cutting repetitive sequences.
Embodiment 3
[0086] Using the pre-sgRNA tandem MICR-CUT system to specifically kill Hela cells.
[0087] In order to enhance the killing effect of the MICR-CUT system, this study chose to connect two pre-sgRNAs targeting different sites in series to form a single pre-sgRNA, so that when the miRNA activates the pre-sgRNA, it can simultaneously produce two different sgRNAs, respectively targeting different genomic repeat sites. In this study, miR-294 and miR-21 targeted activation pre-sgRNAs (referred to as 21T-Alu1-Alu2; 294T-Alu1-Alu2 for short) were constructed.
[0088] See the experimental results image 3 , it can be seen that: after transfection of Cas9 and 21T-Alu1-Alu2 expression plasmids, the number of Hela cells was significantly reduced; while transfection of Cas9 and 294T-Alu1-Alu2 had no significant effect on the growth of Hela cells, only when exogenous transfection of miR -294 can significantly reduce the number of Hela cells. The experimental results further prove that th...
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