Human augmenter of liver regeneration mutant, gene and application
A technology for enhancing factors and liver regeneration, applied in application, genetic engineering, plant genetic improvement, etc., can solve the problems of protein folding and post-translational modification defects, poor rhALR activity, non-uniformity, etc., and achieve improved sulfhydryl oxidation activity and uniform product. , the effect of improving the protection effect
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Embodiment 1
[0036] Referring to the amino acid sequence (81-205 amino acids, a total of 125 amino acids) contained in CAB87993 (http: / / www.ncbi.nlm.nih.gov / protein / cab87993) in Genebank, the codon was optimized and entrusted by Sangon Bioengineering Shanghai The company synthesized the hALR gene (SEQ ID No.3) suitable for Pichia pastoris system expression, cloned it into pPICZα, and constructed the expression plasmid pPICZα-hALR; Anti-transformants, test tube expression screening to obtain expression strains.
[0037] BSM medium was used for fermentation, methanol induction was started when the OD reached above 6, and after 72-96 hours of induction, the tank was placed, and the supernatant was collected by centrifugation.
[0038]The rhALR expressed by yeast is chimeric with FAD molecules during the expression process, and the expression product is a natural dimer. The purified protein solution is transparent yellow, indicating that a stable protein-FAD complex has been formed during the ...
Embodiment 2
[0043] Pichia pastoris secretes and expresses recombinant protein outside the cell, which relies on Kex2 enzyme to recognize and cut KR ↓ (lysine-arginine) sequence. Therefore, when Pichia pastoris expresses the truncated hALR of 125aa, the expression product will be uneven, and most of them will be cleaved by the enzyme to cut off the 7 amino acids at the N-terminus. However, the study found that the deletion of the first 7 amino acids had no effect on its activity. Therefore, mutants missing the first 7 amino acids have been reported in the prior art.
[0044] Considering that the 71st and 72nd amino acid residues are also KR sequences, there may also be the possibility of being recognized and cut by the Kex2 enzyme in Pichia pastoris, so in the present invention, the 71st lysine is mutated into arginine (K71R), the 72nd arginine is mutated to lysine (R72K).
[0045] The natural hALR dimer is a homodimer formed by the reverse combination of two pairs of disulfide bonds at...
Embodiment 3
[0052] Oxidoreductase activity assay. According to the literature (Lisowsky T. et al, Digest Liver Dis., 2001, 33:173) method.
[0053] Accurately weigh glutathione (GSH) and prepare 10mM GSH mother solution with 20mM PB (pH7.5), and accurately weigh DTNB (5,5'-dithiobis(2-nitrobenzoic acid)) to prepare with water into a 10 mM solution; yeast expressed purified rhALR, rhALR (Δ1-7) (prepared in Example 1) and rhALRs (prepared in Example 2), also diluted to 1 mg / mL with 20 mM PB (pH 7.5).
[0054] Prepare the reaction system according to Table 1.
[0055] Table 1 ALR sulfhydryl oxidase activity reaction assay
[0056]
[0057] Leave to react at 30°C for 6 hours, add 20 μL of DTNB to a final concentration of 20 μM, mix thoroughly, continue to react at 30°C for 5 minutes, and measure the absorbance at 412 nm. The sulfhydryl content is calculated according to the formula: sulfhydryl (μM)=OD 412 / 13.6.
[0058] Definition of ALR activity: reaction at 30° C. for 6 hours can r...
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