The invention discloses a longitudinal-axial-flow combine harvester and belongs to the technical field of agricultural machines. The longitudinal-axial-flow combine harvester comprises a harvesting table, a gap bridge device, a threshing separating device, a cleaning device, a grain collecting device and a barn. The threshing separating device comprises a threshing machine body and a longitudinal-axial-flow roller driven by a power device. The longitudinal-axial-flow roller comprises a feeding-in part and a roller body part, and the roller body part sequentially comprises a threshing section, a separating section and a weed-discharging section. A feeding-in shell is arranged on the portion, corresponding to the feeding-in part, of the threshing machine body, a threshing section roller cover is arranged on the upper portion, corresponding to the threshing section, of the threshing machine body, and a threshing section concave plate is arranged on the lower portion, corresponding to the threshing section, of the threshing section roller cover. A separating section roller cover is arranged on the upper portion, corresponding to the separating section, of the threshing machine body, and a separating section concave plate is arranged on the lower portion, corresponding to the separating section, of the threshing machine body. The diameter of the separating section roller cover is larger than that of the threshing section roller cover. The feed quantity is large in harvesting, the function of returning stalks to the field is achieved, working efficiency is improved, kinds of crops are harvested, and the requirements for earnings of a user are met.